Draba incerta |
Draba arabisans |
|
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whitlow-wort, Yellowstone Draba, Yellowstone Draba whitlow-wort, Yellowstone whitlow-grass, Yellowstone whitlow-wort |
rock Draba, rock whitlow-mustard |
|
Habit | Perennials; (cespitose, often pulvinate); caudex branched (dense with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (sometimes cespitose); caudex simple or branched; not scapose. |
Stems | unbranched, (0.2–)0.4–1.4(–2.1) dm, often pubescent throughout, sometimes glabrous distally, trichomes often simple and 2–5-rayed, 0.1–0.5 mm, (sometimes with mostly subpectinate ones). |
branched or unbranched, (0.5–)0.7–3(–4.5) dm, pubescent proximally, glabrous or pubescent distally, trichomes sessile (non-crisped), 4- or 5-rayed, stellate, or subpectinate, 0.07–0.4 mm. |
Basal leaves | rosulate; petiolate; petiole (0–1 cm), ciliate throughout; blade narrowly oblanceolate to linear, (0.4–)0.6–1.7(–2.5) cm × (1–)1.5–3.5(–5) mm, margins entire, (ciliate, trichomes usually simple, rarely 2-rayed, 0.2–1.1 mm), surfaces usually pubescent with short-stalked, pectinate trichomes, 0.15–0.5 mm, sometimes also with 4–6-rayed ones, (midvein usually obscure abaxially), sometimes glabrous adaxially. |
rosulate; petiolate; petiole base and margin sparsely ciliate, (trichomes simple and 2-rayed, 0.2–0.8 mm); blade oblanceolate to spatulate or linear-oblanceolate, 1–3.8(–5) cm × 2–5(–8) mm, margins dentate or denticulate, surfaces pubescent with sessile, non-crisped, 4- or 5-rayed, stellate trichomes, 0.07–0.5mm, (some rays branched, appearing 2–12-rayed, midvein obscure abaxially). |
Cauline leaves | usually 0 (or 1, as a bract); sessile; blade linear to oblong, margins entire, surfaces pubescent as basal. |
(2 or) 3–10(–13); sessile; blade ovate to oblong, margins dentate to subentire, surfaces pubescent as basal. |
Racemes | 3–14(–30)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
5–18(–25)-flowered, ebracteate, elongated in fruit; rachis not flexuous, glabrous or pubescent as stem. |
Flowers | sepals broadly ovate, 2.5–3.5(–4) mm, pubescent, (trichomes simple and 2- or 3-rayed); petals yellow (fading white), oblanceolate to obovate, 4–6 × 1.5–2.5 mm; anthers ovate, 0.3–0.5 mm. |
sepals oblong, 1.7–2.5 mm, pubescent, trichomes short-stalked, 3- or 4-rayed; petals white, broadly obovate, 3.5–5 × 1.5–2.5 mm; anthers ovate, 0.4–0.5 mm. |
Fruiting pedicels | ascending, straight, (2.5–)4–11(–27) mm, glabrous or pubescent, trichomes 2–5-rayed or pectinate. |
divaricate-ascending to suberect, straight or slightly curved, (3–)4–10(–13) mm, glabrous or pubescent as stem. |
Fruits | broadly ovate to lanceolate, plane, flattened, 5–9(–11) × 2–4 mm; valves glabrous or puberulent, trichomes simple and 2-rayed, 0.05–0.3 mm; ovules 8–16(–20) per ovary; style 0.2–0.9 mm. |
ovate to lanceolate or linear-lanceolate, usually slightly twisted, rarely plane, flattened, 5–11(–14) × 2–3.5 mm; valves usually glabrous, rarely pubescent, trichomes 3- or 4-rayed, 0.07–0.2 mm; ovules 20–28(–32) per ovary; style (0.2–)0.4–1 mm. |
Seeds | oblong, 1.1–1.5 × 0.7–1 mm. |
oblong, 0.8–1.1 × 0.6–0.8 mm. |
2n | = 112. |
= 96. |
Draba incerta |
Draba arabisans |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Aug. |
Habitat | Rock outcrops, talus, gravelly areas, tundra | Rock outcrops and talus (usually limestone or dolomite), open woods, rocky shores |
Elevation | 0-3300 m (0-10800 ft) | 0-300 m (0-1000 ft) |
Distribution |
AK; CO; ID; MT; NV; UT; WA; WY; AB; BC; QC; YT
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ME; MI; MN; NY; VT; WI; MB; NB; NF; NS; ON; QC
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Discussion | Draba incerta was shown by G. A. Mulligan (1972) to be sexually reproducing and 14-ploid with x = 8. It is often confused with the apomict D. oligosperma (2n = 32, 64). Draba incerta is readily separated from D. oligosperma by having well-formed (versus abortive) anthers and pollen, stalked (versus sessile) leaf trichomes, and ciliate (versus non-ciliate) basal leaves with obscure (versus prominent) midveins. Although both species have leafless scapes, one often finds a bract adnate to, or subtending, the proximalmost pedicel in D. incerta. Draba incerta is found near sea level in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba arabisans is often confused with D. glabella (2n = 80); it is easily distinguished by having sessile (versus short-stalked) stellate trichomes on abaxial leaf blade surfaces and longer styles (0.2–)0.4–1 [versus 0.1–0.2(–0.4)] mm. Both species have twisted and plane fruits; D. arabisans has mostly twisted fruits; D. glabella has predominantly plane fruits. The complete absence of simple trichomes proximally on stems of D. arabisans can be a useful feature separating it from D. glabella, which often has at least some simple trichomes on the proximal half of the stem. Draba arabis Persoon is an illegitimate name, sometimes found in synonymy with D. arabisans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 313. | FNA vol. 7, p. 291. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. exalata, D. incerta var. laevicapsula, D. incerta var. peasei, D. laevicapsula, D. peasei | D. arabisans var. superiorensis, D. incana var. arabisans, D. incana var. glabriuscula |
Name authority | Payson: Amer. J. Bot. 4: 261. (1917) | Michaux: Fl. Bor.-Amer. 2: 28. (1803) |
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