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twisted Draba

Santa Rita Mountain Draba, Santa Rita Mountain whitlowgrass, Santa Rita whitlow grass

Habit Biennials; not scapose. Perennials; (cespitose, long-lived); caudex simple or branched (covered with persistent petioles), not scapose.
Stems

usually branched, rarely unbranched, (0.3–)0.6–3(–4) dm, densely pubescent proximally, trichomes simple, 0.4–1.3 mm, with fewer 2–4-rayed ones, to 0.3 mm, (distally with mostly branched trichomes, with fewer, simple ones).

usually unbranched, rarely branched, 0.3–1.9(–2.8) dm, moderately to densely hirsute throughout, trichomes simple, 0.2–1.3 mm, with short-stalked to subsessile, cruciform, 0.02–0.4 mm, and 2-rayed ones, 0.3–0.9 mm.

Basal leaves

rosulate;

petiolate;

petiole base and margin ciliate, (trichomes simple, 0.2–0.7 mm);

blade lanceolate or oblanceolate to spatulate, 0.5–2.9 cm × 1–5(–9) mm, margins dentate, surfaces pubescent, abaxially with stalked, cruciform, or 2- or 3-rayed trichomes, 0.1–0.4 mm, (rarely with a branched ray), adaxially with simple trichomes, 0.1–0.7 mm, and stalked, 2–4-rayed ones.

rosulate;

petiolate;

petiole ciliate, (trichomes simple, 0.4–1.5 mm);

blade oblanceolate, 1–5(–6) cm × 2–5(–10) mm, margins usually entire, rarely denticulate, surfaces pubescent, abaxially with stalked, cruciform trichomes, 0.07–0.5 mm, adaxially often similar, sometimes with fewer, simple and 2-rayed trichomes, 0.4–1.3 mm.

Cauline leaves

(15–)22–54(–76), (densely imbricate at least basally);

sessile;

blade ovate to oblong or lanceolate (base often densely bearded abaxially, trichomes simple, to 1.3 mm), margins dentate, surfaces pubescent as basal.

3–10;

sessile;

blade ovate to lanceolate or oblong, margins entire or denticulate, surfaces pubescent as basal.

Racemes

(11–)17–36(–48)-flowered, bracteate basally, elongated or not in fruit;

rachis not flexuous, pubescent as stems distally.

10–37(–58)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

Flowers

sepals ovate, 2–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed);

petals white, spatulate, 3.5–4.5 × 1–2 mm;

anthers ovate, 0.3–0.4 mm.

sepals oblong, 2–3.5 mm, pubescent, (trichomes simple and 2-rayed);

petals yellow, oblanceolate, 3.5–6 × 1.2–1.8 mm;

anthers oblong, 0.8–1 mm.

Fruiting pedicels

erect to ascending, (subappressed to rachis), straight, (1.5–)2.5–6(–13) mm, pubescent, trichomes 2–4-rayed and simple.

divaricate-ascending, straight, 4–10 mm, pubescent throughout, trichomes simple (0.1–0.5 mm) and subsessile, 2–4-rayed, (0.03–0.2 mm).

Fruits

oblong to linear-lanceolate or elliptic, slightly twisted or plane, flattened, 5–10(–12) × 1.5–2.5(–3) mm;

valves glabrous or pubescent, trichomes simple and short-stalked, 2–4-rayed, 0.05–0.3 mm;

ovules (24–)30–40 per ovary;

style 0.01–0.25 mm.

lanceolate to elliptic, often strongly twisted, flattened, 5–11 × 2–3 mm;

valves puberulent at least along margin, trichomes simple, antrorse, 0.03–0.15 mm;

ovules 14–24 per ovary;

style 0.8–1.8(–2.5) mm.

Seeds

oblong, 0.7–1.1 × 0.5–0.6 mm.

ovoid, 1–1.4 × 0.6–0.9 mm.

2n

= 32.

Draba incana

Draba petrophila

Phenology Flowering Jul–Aug. Flowering Jul–Sep.
Habitat Rock outcrops, talus, gravelly shores Crevices, ledges of cliffs
Elevation 0-200 m (0-700 ft) 1200-2800 m (3900-9200 ft)
Distribution
from FNA
MI; MN; MB; NB; NL; ON; PE; QC; Greenland; n Europe; c Europe (Russia)
[BONAP county map]
from FNA
AZ
[WildflowerSearch map]
[BONAP county map]
Discussion

Draba petrophila is often broadly circumscribed to include the taxon herein called D. viridis. For a discussion of the differences between these species and the closely related D. helleriana, see I. A. Al-Shehbaz and M. D. Windham (2007). Draba petrophila, in the strict sense, is known only from Cochise, Pima, and Santa Cruz counties in southeastern Arizona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 312. FNA vol. 7, p. 329.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. confusa, D. contorta, D. incana subsp. confusa, D. incana var. confusa, D. incana var. contorta D. helleriana var. blumeri, D. helleriana var. petrophila
Name authority Linnaeus: Sp. Pl. 2: 643. (1753) Greene: Pittonia 4: 17. (1899)
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