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twisted Draba

schell creek Draba

Habit Biennials; not scapose. Perennials; (cespitose, often forming mats); caudex branched (elongated, with persistent leaf bases, branches sometimes terminating in sterile rosettes); sometimes scapose.
Stems

usually branched, rarely unbranched, (0.3–)0.6–3(–4) dm, densely pubescent proximally, trichomes simple, 0.4–1.3 mm, with fewer 2–4-rayed ones, to 0.3 mm, (distally with mostly branched trichomes, with fewer, simple ones).

unbranched, (0.15–)0.3–0.7(–1) dm, pubescent proximally, trichomes simple and spurred, 0.4–0.8 mm, and short-stalked, 2–4-rayed, 0.1–0.4 mm, (sometimes with simple trichomes distally).

Basal leaves

rosulate;

petiolate;

petiole base and margin ciliate, (trichomes simple, 0.2–0.7 mm);

blade lanceolate or oblanceolate to spatulate, 0.5–2.9 cm × 1–5(–9) mm, margins dentate, surfaces pubescent, abaxially with stalked, cruciform, or 2- or 3-rayed trichomes, 0.1–0.4 mm, (rarely with a branched ray), adaxially with simple trichomes, 0.1–0.7 mm, and stalked, 2–4-rayed ones.

subrosulate;

sessile;

blade oblanceolate to oblong, (0.3–)0.5–0.8 cm × 1–3 mm, (base ciliate, trichomes simple, 0.2–0.8 mm), margins entire, (not ciliate), surfaces usually pubescent, with stalked, (2–)4 (or 5)-rayed trichomes, 0.2–0.5 mm, adaxially sometimes glabrate, or primarily with simple trichomes, 0.3–0.8 mm.

Cauline leaves

(15–)22–54(–76), (densely imbricate at least basally);

sessile;

blade ovate to oblong or lanceolate (base often densely bearded abaxially, trichomes simple, to 1.3 mm), margins dentate, surfaces pubescent as basal.

(0 or) 1–4;

sessile;

blade ovate to oblong, margins entire, surfaces pubescent as basal.

Racemes

(11–)17–36(–48)-flowered, bracteate basally, elongated or not in fruit;

rachis not flexuous, pubescent as stems distally.

(3–)7–15(–20)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

Flowers

sepals ovate, 2–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed);

petals white, spatulate, 3.5–4.5 × 1–2 mm;

anthers ovate, 0.3–0.4 mm.

sepals ovate to oblong, 2–3 mm, pubescent, (trichomes simple, with fewer, short-stalked, 2-rayed ones);

petals white, obovate, 3.5–6 × 1.5–2.8 mm;

anthers oblong, 0.5–0.6 mm.

Fruiting pedicels

erect to ascending, (subappressed to rachis), straight, (1.5–)2.5–6(–13) mm, pubescent, trichomes 2–4-rayed and simple.

divaricate-ascending to ascending, straight, 3–10 mm, pubescent, trichomes simple and short-stalked, 2–4-rayed, (0.1–0.4 mm).

Fruits

oblong to linear-lanceolate or elliptic, slightly twisted or plane, flattened, 5–10(–12) × 1.5–2.5(–3) mm;

valves glabrous or pubescent, trichomes simple and short-stalked, 2–4-rayed, 0.05–0.3 mm;

ovules (24–)30–40 per ovary;

style 0.01–0.25 mm.

broadly or, rarely, narrowly lanceolate to elliptic or ovate, plane or slightly twisted, flattened, (4)5–8(–10) × 2–3.2 mm;

valves (each with distinct midvein), pubescent, trichomes simple and short-stalked, 2-rayed, 0.08–0.2 mm;

ovules 12–24 per ovary;

style (0.7–)1–1.8(–2.1) mm.

Seeds

oblong, 0.7–1.1 × 0.5–0.6 mm.

oblong, 0.9–1.3 × 0.7–1 mm.

2n

= 32.

= 32.

Draba incana

Draba pennellii

Phenology Flowering Jul–Aug. Flowering May–Jul.
Habitat Rock outcrops, talus, gravelly shores Rock outcrops and talus slopes in pinyon-juniper, sagebrush, mountain shrub, and mixed conifer communities
Elevation 0-200 m (0-700 ft) 1900-3500 m (6200-11500 ft)
Distribution
from FNA
MI; MN; MB; NB; NL; ON; PE; QC; Greenland; n Europe; c Europe (Russia)
[BONAP county map]
from FNA
NV
[BONAP county map]
Discussion

Of conservation concern.

Draba pennellii is known from White Pine County in east-central Nevada. It is easily distinguished from other white-flowered species in the region by its elongated, many-branched caudices, long styles, usually few-leaved or, rarely, leafless flowering stems, and distinct midvein on fruit valves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 312. FNA vol. 7, p. 329.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. confusa, D. contorta, D. incana subsp. confusa, D. incana var. confusa, D. incana var. contorta
Name authority Linnaeus: Sp. Pl. 2: 643. (1753) Rollins: J. Arnold Arbor. 64: 502. (1983)
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