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lost river Draba

Draba, tall Draba, tall whitlow-grass, tall whitlow-grass Draba

Habit Perennials; (densely cespitose); caudex branched (densely covered with persistent leaves, branches sometimes terminating in sterile rosettes); scapose. Annuals or perennials; (short-lived); caudex simple or branched; not scapose.
Stems

unbranched, (0.1–)0.3–1(–1.3) dm, hirsute throughout, trichomes simple, 0.4–1 mm, and 2–4-rayed, 0.1–0.6 mm.

branched or unbranched, (0.5–)0.8–3.2(–3.8) dm, pubescent throughout, trichomes 3–5-rayed, 0.1–0.4 mm, with simple and 2-rayed ones, 0.4–1 mm.

Basal leaves

(densely imbricate); rosulate;

sessile;

blade narrowly oblanceolate to oblong-linear, 0.3–1.2(–1.5) cm × 1–2 mm, margins entire, (ciliate, trichomes simple, 0.3–1.2 mm), surfaces pubescent abaxially with stalked, 2–4-rayed trichomes, 0.08–0.45 mm, (midvein obscure), adaxially glabrous proximally, sparsely pubescent distally with mostly simple trichomes.

rosulate;

subsessile;

petiole base ciliate, (trichomes simple and 2-rayed, 0.2–0.8 mm);

blade oblanceolate, (0.7–)1–3.5(–4.5) cm × 1.5–7(–9) mm, margins usually dentate, rarely entire, surfaces pubescent, abaxially with stalked, 3–6-rayed trichomes, 0.1–0.5 mm, adaxially with stalked, 3–5-rayed, 0.1–0.6 mm, and simple ones, to 0.8 mm.

Cauline leaves

0.

(1 or) 2–5(–9);

sessile;

blade ovate to lanceolate, margins dentate or entire, surfaces pubescent as basal.

Racemes

4–15-flowered, ebracteate, elongated in fruit;

rachis not flexuous, hirsute as stem.

(5–)8–30(–37)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

Flowers

sepals oblong, 2.5–3.5 mm, hirsute, (trichomes simple and stalked, 2–4-rayed);

petals white, obovate, 5–6.5 × 2–3.5 mm;

anthers ovate, 0.5–0.6 mm.

(late-season ones petaliferous);

sepals ovate, 1.7–2.5 mm, pubescent, (trichomes simple and stalked, 2-rayed);

petals white, oblanceolate to spatulate, 2.8–3.5(–4) × 0.8–1.2 mm;

anthers ovate, 0.2–0.3 mm.

Fruiting pedicels

ascending, straight, (2–)4–13(–18) mm, hirsute as stem.

divaricate-ascending to ascending or suberect (not appressed to rachis), straight, (3–)4–10(–12) mm, pubescent, trichomes 3–5-rayed (0.1–0.4 mm), and sometimes simple.

Fruits

ovate to broadly oblong or elliptic, plane, flattened, (3–)4–7(–10) × (2.5–)3.5–5 mm;

valves pubescent, trichomes simple and short-stalked, 2(–4)-rayed, 0.06–0.5 mm;

ovules 8–12 per ovary;

style (0.8–)1–1.7(–2) mm.

(erect, not appressed to rachis) lanceolate to linear-lanceolate, plane, flattened, (5–)7–12(–15) × 1.5–2.5(–3) mm;

valves puberulent, trichomes simple and short-stalked, 2–4-rayed, 0.05–0.25 mm;

ovules (24–)30–52 per ovary;

style 0.03–0.1(0.15) mm.

Seeds

oblong, 1.4–1.8 × 0.8–1 mm.

oblong, 0.8–1.1 × 0.5–0.6 mm, (often apiculate).

2n

= 54.

= 56.

Draba hitchcockii

Draba praealta

Phenology Flowering May–Jun. Flowering Jun–Aug.
Habitat Limestone outcrops and gravelly soil Rocky pine slopes, subalpine areas, alpine meadows, grassy volcanic ridges, woodlands, damp rocky areas, limestone talus, shale cliffs, steep hillsides
Elevation 1800-2200 m (5900-7200 ft) 1200-2900 m (3900-9500 ft)
Distribution
from FNA
ID
[BONAP county map]
from FNA
CA; ID; MT; NV; OR; WA; WY; AB; BC; NT; YT; e Asia (Russian Far East [Chukchi Peninsula])
[WildflowerSearch map]
[BONAP county map]
Discussion

Draba hitchcockii is known from the Lost River Range in Butte and Custer counties. Based on morphological and chromosomal evidence, M. D. Windham (2004) suggested that it may be an allopolyploid resulting from hybridization between D. oreibata and D. paysonii. Draba hitchcockii is superficially similar to D. jaegeri, a taxon known from the Charleston Mountains of Clark County, Nevada. Both are cespitose perennials with relatively large, white flowers and a chromosome number (2n = 54) otherwise unknown in Draba (Windham). Features distinguishing these two taxa are provided in the discussion of 51. D. jaegeri.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Morphological and chromosomal evidence (M. D. Windham, unpubl.) suggests that Draba praealta may be an allopolyploid originating through hybridization between D. albertina and D. cana. Both R. C. Rollins (1993) and N. H. Holmgren (2005b) placed D. lonchocarpa var. dasycarpa in the synonymy of D. lonchocarpa. G. A. Mulligan (1974) indicated that the type of var. dasycarpa does not belong to that species and he correctly annotated its isotype (Macoun 64454, CAN) as D. praealta. Mulligan (1976) and Rollins indicated that D. praealta occurs in Alaska, but we have not seen unequivocal material from the state.

We have not seen the material on which A. E. Katenin and V. V. Petrovsky (1995) based their record from the Russian Far East.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 311. FNA vol. 7, p. 331.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. cascadensis, D. columbiana, D. dolichopoda, D. lapilutea, D. lonchocarpa var. dasycarpa, D. praealta var. yellowstonensis, D. yellowstonensis
Name authority Rollins: J. Arnold Arbor. 64: 500. (1983) Greene: Pittonia 3: 306. (1898)
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