Draba hitchcockii |
Draba praealta |
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lost river Draba |
Draba, tall Draba, tall whitlow-grass, tall whitlow-grass Draba |
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Habit | Perennials; (densely cespitose); caudex branched (densely covered with persistent leaves, branches sometimes terminating in sterile rosettes); scapose. | Annuals or perennials; (short-lived); caudex simple or branched; not scapose. |
Stems | unbranched, (0.1–)0.3–1(–1.3) dm, hirsute throughout, trichomes simple, 0.4–1 mm, and 2–4-rayed, 0.1–0.6 mm. |
branched or unbranched, (0.5–)0.8–3.2(–3.8) dm, pubescent throughout, trichomes 3–5-rayed, 0.1–0.4 mm, with simple and 2-rayed ones, 0.4–1 mm. |
Basal leaves | (densely imbricate); rosulate; sessile; blade narrowly oblanceolate to oblong-linear, 0.3–1.2(–1.5) cm × 1–2 mm, margins entire, (ciliate, trichomes simple, 0.3–1.2 mm), surfaces pubescent abaxially with stalked, 2–4-rayed trichomes, 0.08–0.45 mm, (midvein obscure), adaxially glabrous proximally, sparsely pubescent distally with mostly simple trichomes. |
rosulate; subsessile; petiole base ciliate, (trichomes simple and 2-rayed, 0.2–0.8 mm); blade oblanceolate, (0.7–)1–3.5(–4.5) cm × 1.5–7(–9) mm, margins usually dentate, rarely entire, surfaces pubescent, abaxially with stalked, 3–6-rayed trichomes, 0.1–0.5 mm, adaxially with stalked, 3–5-rayed, 0.1–0.6 mm, and simple ones, to 0.8 mm. |
Cauline leaves | 0. |
(1 or) 2–5(–9); sessile; blade ovate to lanceolate, margins dentate or entire, surfaces pubescent as basal. |
Racemes | 4–15-flowered, ebracteate, elongated in fruit; rachis not flexuous, hirsute as stem. |
(5–)8–30(–37)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals oblong, 2.5–3.5 mm, hirsute, (trichomes simple and stalked, 2–4-rayed); petals white, obovate, 5–6.5 × 2–3.5 mm; anthers ovate, 0.5–0.6 mm. |
(late-season ones petaliferous); sepals ovate, 1.7–2.5 mm, pubescent, (trichomes simple and stalked, 2-rayed); petals white, oblanceolate to spatulate, 2.8–3.5(–4) × 0.8–1.2 mm; anthers ovate, 0.2–0.3 mm. |
Fruiting pedicels | ascending, straight, (2–)4–13(–18) mm, hirsute as stem. |
divaricate-ascending to ascending or suberect (not appressed to rachis), straight, (3–)4–10(–12) mm, pubescent, trichomes 3–5-rayed (0.1–0.4 mm), and sometimes simple. |
Fruits | ovate to broadly oblong or elliptic, plane, flattened, (3–)4–7(–10) × (2.5–)3.5–5 mm; valves pubescent, trichomes simple and short-stalked, 2(–4)-rayed, 0.06–0.5 mm; ovules 8–12 per ovary; style (0.8–)1–1.7(–2) mm. |
(erect, not appressed to rachis) lanceolate to linear-lanceolate, plane, flattened, (5–)7–12(–15) × 1.5–2.5(–3) mm; valves puberulent, trichomes simple and short-stalked, 2–4-rayed, 0.05–0.25 mm; ovules (24–)30–52 per ovary; style 0.03–0.1(0.15) mm. |
Seeds | oblong, 1.4–1.8 × 0.8–1 mm. |
oblong, 0.8–1.1 × 0.5–0.6 mm, (often apiculate). |
2n | = 54. |
= 56. |
Draba hitchcockii |
Draba praealta |
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Phenology | Flowering May–Jun. | Flowering Jun–Aug. |
Habitat | Limestone outcrops and gravelly soil | Rocky pine slopes, subalpine areas, alpine meadows, grassy volcanic ridges, woodlands, damp rocky areas, limestone talus, shale cliffs, steep hillsides |
Elevation | 1800-2200 m (5900-7200 ft) | 1200-2900 m (3900-9500 ft) |
Distribution |
ID |
CA; ID; MT; NV; OR; WA; WY; AB; BC; NT; YT; e Asia (Russian Far East [Chukchi Peninsula])
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Discussion | Draba hitchcockii is known from the Lost River Range in Butte and Custer counties. Based on morphological and chromosomal evidence, M. D. Windham (2004) suggested that it may be an allopolyploid resulting from hybridization between D. oreibata and D. paysonii. Draba hitchcockii is superficially similar to D. jaegeri, a taxon known from the Charleston Mountains of Clark County, Nevada. Both are cespitose perennials with relatively large, white flowers and a chromosome number (2n = 54) otherwise unknown in Draba (Windham). Features distinguishing these two taxa are provided in the discussion of 51. D. jaegeri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological and chromosomal evidence (M. D. Windham, unpubl.) suggests that Draba praealta may be an allopolyploid originating through hybridization between D. albertina and D. cana. Both R. C. Rollins (1993) and N. H. Holmgren (2005b) placed D. lonchocarpa var. dasycarpa in the synonymy of D. lonchocarpa. G. A. Mulligan (1974) indicated that the type of var. dasycarpa does not belong to that species and he correctly annotated its isotype (Macoun 64454, CAN) as D. praealta. Mulligan (1976) and Rollins indicated that D. praealta occurs in Alaska, but we have not seen unequivocal material from the state. We have not seen the material on which A. E. Katenin and V. V. Petrovsky (1995) based their record from the Russian Far East. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 311. | FNA vol. 7, p. 331. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. cascadensis, D. columbiana, D. dolichopoda, D. lapilutea, D. lonchocarpa var. dasycarpa, D. praealta var. yellowstonensis, D. yellowstonensis | |
Name authority | Rollins: J. Arnold Arbor. 64: 500. (1983) | Greene: Pittonia 3: 306. (1898) |
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