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lost river Draba

palander's Draba, palander's whitlow-grass

Habit Perennials; (densely cespitose); caudex branched (densely covered with persistent leaves, branches sometimes terminating in sterile rosettes); scapose. Perennials; (cespitose); caudex branched (with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose.
Stems

unbranched, (0.1–)0.3–1(–1.3) dm, hirsute throughout, trichomes simple, 0.4–1 mm, and 2–4-rayed, 0.1–0.6 mm.

unbranched, 0.2–1.1(–1.5) dm, pubescent proximally, trichomes 2–9-rayed, 0.05–0.2 mm, glabrous or sparsely pubescent distally.

Basal leaves

(densely imbricate); rosulate;

sessile;

blade narrowly oblanceolate to oblong-linear, 0.3–1.2(–1.5) cm × 1–2 mm, margins entire, (ciliate, trichomes simple, 0.3–1.2 mm), surfaces pubescent abaxially with stalked, 2–4-rayed trichomes, 0.08–0.45 mm, (midvein obscure), adaxially glabrous proximally, sparsely pubescent distally with mostly simple trichomes.

rosulate;

petiolate;

petiole (thickened), ciliate proximally, (margin not ciliate, trichomes simple and 2-rayed, 0.2–0.5 mm);

blade oblanceolate to obovate, 0.3–0.9(–1.5) cm × 1–4 mm, margins usually entire, rarely denticulate, surfaces pubescent with minutely stalked, 8–16-rayed, stellate trichomes, 0.08–0.3 mm, (midvein obscure abaxially).

Cauline leaves

0.

0.

Racemes

4–15-flowered, ebracteate, elongated in fruit;

rachis not flexuous, hirsute as stem.

5–17-flowered, ebracteate, elongated in fruit;

rachis not flexuous, usually glabrous, rarely sparsely pubescent as stem.

Flowers

sepals oblong, 2.5–3.5 mm, hirsute, (trichomes simple and stalked, 2–4-rayed);

petals white, obovate, 5–6.5 × 2–3.5 mm;

anthers ovate, 0.5–0.6 mm.

sepals ovate, 2–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–6-rayed);

petals white or creamy white, obovate, 4.5–5.5 × 2–3 mm;

anthers ovate, 0.4–0.6 mm.

Fruiting pedicels

ascending, straight, (2–)4–13(–18) mm, hirsute as stem.

divaricate-ascending, straight, (slender), 4–10 mm, usually glabrous, rarely sparsely pubescent as stem.

Fruits

ovate to broadly oblong or elliptic, plane, flattened, (3–)4–7(–10) × (2.5–)3.5–5 mm;

valves pubescent, trichomes simple and short-stalked, 2(–4)-rayed, 0.06–0.5 mm;

ovules 8–12 per ovary;

style (0.8–)1–1.7(–2) mm.

(often aborting, becoming lopsided), oblong to elliptic or ovate, plane, flattened, 4–8 × 2–3 mm;

valves glabrous;

ovules 18–32 per ovary;

style 0.3 0.8 mm.

Seeds

oblong, 1.4–1.8 × 0.8–1 mm.

ovoid, 0.9–1.1 × 0.5–0.7 mm.

2n

= 54.

= 16, 32, 64.

Draba hitchcockii

Draba palanderiana

Phenology Flowering May–Jun. Flowering Jun–Jul.
Habitat Limestone outcrops and gravelly soil Rock outcrops, talus, Dryas fellfields, tundra
Elevation 1800-2200 m (5900-7200 ft) 0-1800 m (0-5900 ft)
Distribution
from FNA
ID
[BONAP county map]
from FNA
AK; NT; YT; e Asia (Russian Far East, Siberia)
[BONAP county map]
Discussion

Draba hitchcockii is known from the Lost River Range in Butte and Custer counties. Based on morphological and chromosomal evidence, M. D. Windham (2004) suggested that it may be an allopolyploid resulting from hybridization between D. oreibata and D. paysonii. Draba hitchcockii is superficially similar to D. jaegeri, a taxon known from the Charleston Mountains of Clark County, Nevada. Both are cespitose perennials with relatively large, white flowers and a chromosome number (2n = 54) otherwise unknown in Draba (Windham). Features distinguishing these two taxa are provided in the discussion of 51. D. jaegeri.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

North American plants of Draba palanderiana are diploid and appear to be self-incompatible, an unusual situation among arctic members of the genus. By contrast, some plants from the Russian Far East are tetraploid (2n = 32) or octoploid (2n = 64) and appear to be self-compatible. Detailed studies are needed to establish whether one or two taxa are involved.

Many North American collections of Draba palanderiana have been misidentified as D. nivalis. The species is easily separated from D. nivalis by having ciliate (versus non-ciliate) bases of basal leaves, larger petals (4.5–5.5 × 2–3 versus 2–3.5 × 0.8–1.4 mm) that are pale yellow or cream (versus white), and longer fruiting pedicels 4–10 [versus 1–4.5(–8)] mm.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 311. FNA vol. 7, p. 326.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Name authority Rollins: J. Arnold Arbor. 64: 500. (1983) Kjellman: in A. E. Nordenskiöld, Vega Exp. Vetensk. Iakttag. 2: 45. (1883)
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