Draba hitchcockii |
Draba novolympica |
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lost river Draba |
Mt. Olympic Draba, Olympic Draba, Olympic Mountains Draba, Payson's Draba, Payson's whitlow-grass, Trelease's Draba, Trelease's whitlow-grass |
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Habit | Perennials; (densely cespitose); caudex branched (densely covered with persistent leaves, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (cespitose, densely pulvinate); caudex branched (covered with persistent leaves, branches creeping, sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, (0.1–)0.3–1(–1.3) dm, hirsute throughout, trichomes simple, 0.4–1 mm, and 2–4-rayed, 0.1–0.6 mm. |
unbranched, 0.05–0.4 dm, densely pubescent throughout, trichomes simple, 0.4–0.8 mm, and stalked, 2–5-rayed, 0.1–0.5 mm. |
Basal leaves | (densely imbricate); rosulate; sessile; blade narrowly oblanceolate to oblong-linear, 0.3–1.2(–1.5) cm × 1–2 mm, margins entire, (ciliate, trichomes simple, 0.3–1.2 mm), surfaces pubescent abaxially with stalked, 2–4-rayed trichomes, 0.08–0.45 mm, (midvein obscure), adaxially glabrous proximally, sparsely pubescent distally with mostly simple trichomes. |
(densely imbricate); rosulate; sessile; blade oblong to linear-oblanceolate, 0.2–0.8 cm × 0.5–1.5 mm, margins entire, (ciliate, trichomes simple and spurred, 0.3–1.2 mm), surfaces densely pubescent, abaxially with stalked, 2–12-rayed stellate trichomes, 0.1–0.6 mm, adaxially with simple and 2-rayed ones, 0.3–0.8 mm. |
Cauline leaves | 0. |
0. |
Racemes | 4–15-flowered, ebracteate, elongated in fruit; rachis not flexuous, hirsute as stem. |
2–12-flowered, ebracteate, slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals oblong, 2.5–3.5 mm, hirsute, (trichomes simple and stalked, 2–4-rayed); petals white, obovate, 5–6.5 × 2–3.5 mm; anthers ovate, 0.5–0.6 mm. |
sepals oblong, 1.5–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals bright yellow, oblanceolate to spatulate, 2–3.5(–4) × 1.5–2 mm; anthers oblong, 0.5–0.6 mm. |
Fruiting pedicels | ascending, straight, (2–)4–13(–18) mm, hirsute as stem. |
divaricate-ascending, straight, 1–5 mm, pubescent, trichomes simple (0.3–0.9 mm) and stalked, 2–5-rayed (0.1–0.5 mm). |
Fruits | ovate to broadly oblong or elliptic, plane, flattened, (3–)4–7(–10) × (2.5–)3.5–5 mm; valves pubescent, trichomes simple and short-stalked, 2(–4)-rayed, 0.06–0.5 mm; ovules 8–12 per ovary; style (0.8–)1–1.7(–2) mm. |
often ovoid, plane, slightly inflated basally (symmetric), (2.5–)3–4(–5) × 1.5–3.5 mm; valves densely pubescent, trichomes 2–6-rayed, 0.05–0.4 mm, occasionally some simple; ovules 4–8(–12) per ovary; style 0.2–0.6(–0.8) mm. |
Seeds | oblong, 1.4–1.8 × 0.8–1 mm. |
oblong, 1.2–1.8 × 0.8–1.1 mm. |
2n | = 54. |
= 42. |
Draba hitchcockii |
Draba novolympica |
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Phenology | Flowering May–Jun. | Flowering Jun–Aug. |
Habitat | Limestone outcrops and gravelly soil | Alpine crests, open knolls, fellfields, talus, weathered shale, calcareous shale scree, rocky grounds and cliffs, subalpine conifer forests |
Elevation | 1800-2200 m (5900-7200 ft) | 1500-3700 m (4900-12100 ft) |
Distribution |
ID |
CA; ID; MT; NV; OR; UT; WA; WY; AB; BC
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Discussion | Draba hitchcockii is known from the Lost River Range in Butte and Custer counties. Based on morphological and chromosomal evidence, M. D. Windham (2004) suggested that it may be an allopolyploid resulting from hybridization between D. oreibata and D. paysonii. Draba hitchcockii is superficially similar to D. jaegeri, a taxon known from the Charleston Mountains of Clark County, Nevada. Both are cespitose perennials with relatively large, white flowers and a chromosome number (2n = 54) otherwise unknown in Draba (Windham). Features distinguishing these two taxa are provided in the discussion of 51. D. jaegeri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba novolympica is the same taxon that C. L. Hitchcock (1941) and R. C. Rollins (1993) called D. paysonii var. treleasei, and G. A. Mulligan (2002) called D. paysonii. The two are amply distinct and should be recognized as separate species. Draba novolympica is easily distinguished from D. paysonii by having fruit valves pubescent with 2–6-rayed (occasionally some simple) trichomes 0.05–0.4 mm, sepals 1.5–2.5 mm, petals 2–3.5(–4) × 1.5–2 mm, fruits (2.5–)3–4(–5) × 1.5–3.5 mm, styles 0.2–0.6(–0.8) mm, and ovules 1.2–1.8 × 0.8–1.1 mm. By contrast, D. paysonii has fruit valves pubescent with simple and 2-rayed (some 4- or 5-rayed) trichomes (0.2–)0.4–1 mm, sepals 2.8–3.5 mm, petals (4–)5–6 × (1.5–)2–3 mm, fruits (5–)6–9 × (3–)3.5–5 mm, styles (0.6–)0.8–1.2 mm, and ovules 1.7–2.2 × 1–1.4 mm. Both R. C. Rollins (1993) and N. H. Holmgren (2005b) indicated that Draba novolympica (as D. paysonii var. treleasei) occurs in Alaska and Yukon, but we have not seen any material from there, and it is likely that their records were based on misidentified plants. Previous reports of D. paysonii from Canada (e.g., G. A. Mulligan 1971b) pertain instead to D. novolympica. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 311. | FNA vol. 7, p. 323. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. barbata var. treleasei, D. paysonii var. treleasei | |
Name authority | Rollins: J. Arnold Arbor. 64: 500. (1983) | Payson & H. St. John: Proc. Biol. Soc. Wash. 43: 113. (1930) |
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