Draba helleriana |
Draba yukonensis |
|
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Heller's Draba |
Yukon whitlowgrass |
|
Habit | Biennials or perennials; (short-lived); caudex simple or branched; not scapose. | Perennials; (short-lived); caudex simple or branched (with persistent leaf remains); not scapose. |
Stems | branched, (0.1–)1.5–4.3(–5.1) dm, hirsute throughout, trichomes simple, 0.4–1.3(–1.8) mm, with short-stalked to subsessile, 3–5-rayed ones, 0.2–0.4 mm, (some slightly coarser, 2-rayed). |
branched, 0.4–2 dm, pubescent throughout, trichomes simple, 0.3–1.2 mm, and 2–6-rayed, 0.05–0.6 mm. |
Basal leaves | rosulate; petiolate; petiole base ciliate or not; blade oblanceolate to obovate, 0.9–4.1(–5.2) cm × 2–7(–10) mm, margins entire or dentate, surfaces pubescent, abaxially with stalked, cruciform, and fewer 3–5-rayed trichomes, 0.1–0.6 mm, adaxially with cruciform and simple trichomes, 0.4–1.3 mm, and smaller 2-rayed ones. |
rosulate; subsessile; blade oblanceolate to narrowly so, 0.3–1 cm × 0.5–2 mm, margins entire, (ciliate, trichomes simple, to 1.3 mm), surfaces pubescent, abaxially with (rigid), simple and 2-rayed trichomes, 0.4–1.2 mm, sometimes with short-stalked, 3–6-rayed ones, 0.1–0.3 mm, adaxially with simple trichomes. |
Cauline leaves | (8–)12–31(–43); sessile; blade ovate to lanceolate or oblong, margins usually dentate, rarely subentire, surfaces pubescent as basal. |
1–3; sessile; blade ovate to oblong, margins entire, surfaces pubescent as basal. |
Racemes | 10–52(–83)-flowered, usually ebracteate, rarely proximalmost 1 or 2 flowers bracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
5–20-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals oblong, 2.5–4 mm, pubescent, (trichomes simple and stalked, 2-rayed); petals yellow, oblanceolate, 5–7 × 1.5–2.2 mm; anthers oblong, 0.5–0.8 mm. |
sepals ovate, 1.2–1.6 mm, pubescent, (trichomes simple and 2–4-rayed); petals white, spatulate, 1.5–2 × 0.7–0.8 mm; anthers ovate, ca. 0.2 mm. |
Fruiting pedicels | horizontal to divaricate-ascending, straight or slightly curved upward, 4–10(–13) mm, pubescent as rachis abaxially, glabrous adaxially. |
erect to ascending, straight, 0.5–3 mm, pubescent as stem. |
Fruits | (not appressed to rachis), lanceolate to ovate or oblong-lanceolate, slightly to strongly twisted or plane, flattened, 5–15 × 2–3.5 mm; valves puberulent, trichomes simple and subsessile, 2(–4)-rayed, 0.03–0.25(–0.8) mm; ovules 14–28 per ovary; style (1–)1.5–3.5 mm. |
ovoid to oblong, plane, not flattened, 2–5 × 1.5–2 mm; valves pubescent, trichomes short-stalked, 4- or 5-rayed, 0.05–0.3 mm; ovules 36–52 per ovary; style 0.1–0.3 mm. |
Seeds | oblong, 1–1.3 × 0.6–0.8 mm. |
oblong, 0.6–0.8 × 0.3–0.4 mm. |
2n | = 18. |
|
Draba helleriana |
Draba yukonensis |
|
Phenology | Flowering Jun–Sep. | Flowering late May–Jun. |
Habitat | Oak and pine-fir woodlands, aspen groves, rocky meadows | Open stony ridges on ancient beach in grassy areas and aspen stands |
Elevation | 2100-3600 m (6900-11800 ft) | ca. 600 m (ca. 2000 ft) |
Distribution |
AZ; CO; NM; Mexico (Nuevo León)
|
YT |
Discussion | Draba helleriana is highly variable and was divided by C. L. Hitchcock (1941) into four varieties. For a discussion of those and the circumscription of the species, see I. A. Al-Shehbaz and M. D. Windham (2007). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Draba yukonensis is known only from Kluane National Park, southwestern Yukon Territory. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 311. | FNA vol. 7, p. 346. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. aurea var. tylosa, D. helleriana var. leiocarpa, D. helleriana var. neomexicana, D. helleriana var. patens, D. helleriana var. pinetorum, D. neomexicana, D. neomexicana var. robusta, D. pallida, D. patens, D. pinetorum, D. stylosa | |
Name authority | Greene: Pittonia 4: 17. (1899) | A. E. Porsild: Publ. Bot. (Ottawa) 4: 37, plate 7. (1975) |
Web links |