Flora of North America species comparison

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Draba helleriana

Draba verna

Heller's Draba

common Draba, common whitlow-grass, early witlow grass, nailwort, shadflower, spring Draba, spring whitlow-grass, spring whitlow-mustard, vernal whitlow grass, whitlow grass, whitlow wort
Habit Biennials or perennials; (short-lived); caudex simple or branched; not scapose. Annuals; scapose.
Stems

branched, (0.1–)1.5–4.3(–5.1) dm, hirsute throughout, trichomes simple, 0.4–1.3(–1.8) mm, with short-stalked to subsessile, 3–5-rayed ones, 0.2–0.4 mm, (some slightly coarser, 2-rayed).

(few to many from base), unbranched, (0.2–)0.5–2(–3) dm, pubescent proximally, glabrous distally, trichomes simple and 2(–4)-rayed, 0.1–0.4 mm.
Basal leaves

rosulate;

petiolate;

petiole base ciliate or not;

blade oblanceolate to obovate, 0.9–4.1(–5.2) cm × 2–7(–10) mm, margins entire or dentate, surfaces pubescent, abaxially with stalked, cruciform, and fewer 3–5-rayed trichomes, 0.1–0.6 mm, adaxially with cruciform and simple trichomes, 0.4–1.3 mm, and smaller 2-rayed ones.

rosulate;

petiolate;

blade obovate, spatulate, oblanceolate, lanceolate, oblong, or, rarely, linear, 0.2–1.8(–3) cm × (0.5–)1–5(–10) mm, margins entire or 1–5-toothed on each side, surfaces pubescent with simple or stalked, 2–4-rayed trichomes, 0.1–0.5 mm.
Cauline leaves

(8–)12–31(–43);

sessile;

blade ovate to lanceolate or oblong, margins usually dentate, rarely subentire, surfaces pubescent as basal.

0.
Racemes

10–52(–83)-flowered, usually ebracteate, rarely proximalmost 1 or 2 flowers bracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

4–20(–30)-flowered, ebracteate, usually considerably elongated in fruit;

rachis usually flexuous, glabrous.
Flowers

sepals oblong, 2.5–4 mm, pubescent, (trichomes simple and stalked, 2-rayed);

petals yellow, oblanceolate, 5–7 × 1.5–2.2 mm;

anthers oblong, 0.5–0.8 mm.

sepals (green or purplish), oblong, 1–2.5 mm, glabrescent or pubescent, (trichomes simple or 2-rayed);

petals white, deeply 2-fid, (1.5–)2–4.5(–6) × 1–2 mm;

anthers ovate, 0.2–0.4 mm.
Fruiting pedicels

horizontal to divaricate-ascending, straight or slightly curved upward, 4–10(–13) mm, pubescent as rachis abaxially, glabrous adaxially.

divaricate to ascending, straight or slightly curved upward, (2–)5–20(–35) mm, glabrous.
Fruits

(not appressed to rachis), lanceolate to ovate or oblong-lanceolate, slightly to strongly twisted or plane, flattened, 5–15 × 2–3.5 mm;

valves puberulent, trichomes simple and subsessile, 2(–4)-rayed, 0.03–0.25(–0.8) mm;

ovules 14–28 per ovary;

style (1–)1.5–3.5 mm.

obovate, oblanceolate, lanceolate, elliptic, oblong, or linear, plane, flattened, (2.5–)4–9(–12) × 1.5–2.5(–3.5) mm;

valves glabrous;

ovules (20–)32–70(–84) per ovary;

style 0.02–0.2 mm.
Seeds

oblong, 1–1.3 × 0.6–0.8 mm.

ovoid (slightly flattened), 0.3–0.6(–0.8) × 0.2–0.4 mm.
2n

= 18.

= 14, 16, 20, 24, 28, 30, 32, 34, 36, 38, 40, 52, 54, 58, 60, 64.

Draba helleriana

Draba verna
Phenology Flowering Jun–Sep. Flowering Feb–May.
Habitat Oak and pine-fir woodlands, aspen groves, rocky meadows Cedar glades, lawns, fields, pastures, waste places, grassy hillsides, disturbed sites, roadsides
Elevation 2100-3600 m (6900-11800 ft) 0-2500 m (0-8200 ft)
Distribution
from FNA
AZ; CO; NM; Mexico (Nuevo León)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CA; CT; DC; DE; GA; IA; ID; IL; IN; KY; MA; MD; ME; MI; MO; MS; MT; NC; NH; NJ; NV; NY; OH; OR; PA; RI; SC; TN; UT; VA; VT; WA; WV; WY; AB; BC; NB; ON; QC; Europe; Asia; nw Africa [Introduced in North America; introduced also in Central America, South America, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Draba helleriana is highly variable and was divided by C. L. Hitchcock (1941) into four varieties. For a discussion of those and the circumscription of the species, see I. A. Al-Shehbaz and M. D. Windham (2007).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba verna represents a highly variable and taxonomically difficult complex within which species, subspecies, varieties, and forms have been named (O. E. Schulz 1927); only those synonyms pertaining to North America are listed above. Most of the taxonomic difficulties are the results of disploidy, autogamy, and hybridization. The morphological extremes are connected by intermediate forms in every conceivable character. Furthermore, there appears to be no correlation between morphology, cytology, geography, and ecology to support the division of this complex into meaningful taxa. A complex cytological picture was presented by Ø. Winge (1940), including the highest count of 2n = 94, which has not been confirmed by subsequent botanists.

Erophila vulgaris de Candolle is an illegitimate name for Draba verna.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 7, p. 311. FNA vol. 7, p. 345.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. aurea var. tylosa, D. helleriana var. leiocarpa, D. helleriana var. neomexicana, D. helleriana var. patens, D. helleriana var. pinetorum, D. neomexicana, D. neomexicana var. robusta, D. pallida, D. patens, D. pinetorum, D. stylosa D. boerhaavii, D. praecox, D. verna var. aestivalis, D. verna var. boerhaavii, Erophila boerhaavii, Erophila krockeri, Erophila praecox, Erophila verna, Erophila verna subsp. praecox
Name authority Greene: Pittonia 4: 17. (1899) Linnaeus: Sp. Pl. 2: 642. (1753)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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