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Heller's Draba
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mound Draba, parasol Draba
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| Biennials or perennials; (short-lived); caudex simple or branched; not scapose. |
Perennials; (cespitose, densely pulvinate); caudex branched (densely covered with persistent leaves and remains, branches tightly grouped, sometimes terminating in sterile rosettes); scapose. |
branched, (0.1–)1.5–4.3(–5.1) dm, hirsute throughout, trichomes simple, 0.4–1.3(–1.8) mm, with short-stalked to subsessile, 3–5-rayed ones, 0.2–0.4 mm, (some slightly coarser, 2-rayed). |
unbranched, (0.05–)0.1–0.25 dm, densely pubescent throughout, trichomes (grayish), stalked, dendritic, 5–12-rayed, 0.1–0.25 mm. |
rosulate; petiolate; petiole base ciliate or not; blade oblanceolate to obovate, 0.9–4.1(–5.2) cm × 2–7(–10) mm, margins entire or dentate, surfaces pubescent, abaxially with stalked, cruciform, and fewer 3–5-rayed trichomes, 0.1–0.6 mm, adaxially with cruciform and simple trichomes, 0.4–1.3 mm, and smaller 2-rayed ones. |
(densely imbricate); rosulate; sessile; blade obovate to broadly oblong, 0.2–0.4 cm × 0.6–1.5 mm, margins entire, (not ciliate), surfaces densely pubescent, trichomes (grayish), stalked, dendritic, 5–12-rayed, 0.1–0.3 mm, (sometimes with long-stalked and spurred trichomes adaxially). |
(8–)12–31(–43); sessile; blade ovate to lanceolate or oblong, margins usually dentate, rarely subentire, surfaces pubescent as basal. |
0. |
10–52(–83)-flowered, usually ebracteate, rarely proximalmost 1 or 2 flowers bracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
2–5(–10)-flowered, ebracteate, subumbellate or slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
sepals oblong, 2.5–4 mm, pubescent, (trichomes simple and stalked, 2-rayed); petals yellow, oblanceolate, 5–7 × 1.5–2.2 mm; anthers oblong, 0.5–0.8 mm. |
sepals broadly oblong, 1.8–2.8 mm, pubescent, (trichomes dendritic); petals yellow, spatulate, 2.8–4 × 1–1.5 mm; anthers ovate, 0.3–0.4 mm. |
horizontal to divaricate-ascending, straight or slightly curved upward, 4–10(–13) mm, pubescent as rachis abaxially, glabrous adaxially. |
divaricate-ascending, straight, 1.5–3(–6) mm, glabrous or pubescent as stem. |
(not appressed to rachis), lanceolate to ovate or oblong-lanceolate, slightly to strongly twisted or plane, flattened, 5–15 × 2–3.5 mm; valves puberulent, trichomes simple and subsessile, 2(–4)-rayed, 0.03–0.25(–0.8) mm; ovules 14–28 per ovary; style (1–)1.5–3.5 mm. |
ovoid to ovoid-lanceolate, plane, inflated and sometimes subgibbous basally, flattened distally, 2–5 × 2–3 mm; valves pubescent, trichomes short-stalked, dendritic, 4–12-rayed, (sometimes spurred), 0.05–0.2 mm; ovules 6–12 per ovary; style 0.2–0.6 mm. |
oblong, 1–1.3 × 0.6–0.8 mm. |
oblong, 1–1.2 × 0.5–0.7 mm. |
= 18. |
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| Flowering Jun–Sep. |
Flowering Jul–Aug. |
| Oak and pine-fir woodlands, aspen groves, rocky meadows |
Wind-eroded areas, alpine fellfields, rock crevices |
| 2100-3600 m (6900-11800 ft) |
3300-4100 m (10800-13500 ft) |
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AZ; CO; NM; Mexico (Nuevo León)
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CA; NV |
Draba helleriana is highly variable and was divided by C. L. Hitchcock (1941) into four varieties. For a discussion of those and the circumscription of the species, see I. A. Al-Shehbaz and M. D. Windham (2007). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba subumbellata is similar to, and sympatric with, D. oligosperma. It is distinguished from the latter by having stalked, stellate to dendritic leaf trichomes, subumbellate racemes, fruiting pedicels 1.5–3.0(–6.0) mm, fruits pubescent with mostly 4–12-rayed trichomes, and well-formed anthers and pollen. By contrast, D. oligosperma has sessile or subsessile, pectinately-branched leaf trichomes, elongated racemes, fruiting pedicels (2–)3–10(–13) mm, fruits glabrous or pubescent with simple and 2-rayed trichomes, and abortive anthers and/or pollen. Draba subumbellata is restricted to the White Mountains (Esmeralda County, Nevada, and Mono County, California) and to Coyote Ridge in the Sierra Nevada (Inyo County, California). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 7, p. 311. |
FNA vol. 7, p. 343. |
| Brassicaceae > tribe Arabideae > Draba |
Brassicaceae > tribe Arabideae > Draba |
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis |
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis |
| D. aurea var. tylosa, D. helleriana var. leiocarpa, D. helleriana var. neomexicana, D. helleriana var. patens, D. helleriana var. pinetorum, D. neomexicana, D. neomexicana var. robusta, D. pallida, D. patens, D. pinetorum, D. stylosa |
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| Greene: Pittonia 4: 17. (1899) |
Rollins & R. A. Price: Aliso 12: 25, figs. 1k–n, 4. (1988) |
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