Draba helleriana |
Draba stenoloba |
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Heller's Draba |
Alaska Draba, Alaska whitlow-grass |
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Habit | Biennials or perennials; (short-lived); caudex simple or branched; not scapose. | Biennials or perennials; (short-lived, not cespitose); caudex simple (not fleshy); scapose. |
Stems | branched, (0.1–)1.5–4.3(–5.1) dm, hirsute throughout, trichomes simple, 0.4–1.3(–1.8) mm, with short-stalked to subsessile, 3–5-rayed ones, 0.2–0.4 mm, (some slightly coarser, 2-rayed). |
often unbranched, (0.2–)0.4–3(–3.4) dm, pubescent proximally, usually glabrous distally, rarely sparsely pubescent, trichomes short-stalked, 2–4-rayed, 0.1–0.4(–0.5) mm. |
Basal leaves | rosulate; petiolate; petiole base ciliate or not; blade oblanceolate to obovate, 0.9–4.1(–5.2) cm × 2–7(–10) mm, margins entire or dentate, surfaces pubescent, abaxially with stalked, cruciform, and fewer 3–5-rayed trichomes, 0.1–0.6 mm, adaxially with cruciform and simple trichomes, 0.4–1.3 mm, and smaller 2-rayed ones. |
rosulate; petiole (obscure), margin ciliate, (trichomes simple, 0.2–0.4 mm); blade oblanceolate to obovate, 0.4–2.6(–3.1) cm × 1.5–7(–10) mm, margins entire or denticulate, (ciliate as petiole), surfaces pubescent, abaxially with short-stalked, 3- or 4-rayed trichomes, 0.1–0.5 mm, adaxially with simple and stalked, 2–4-rayed trichomes, 0.1–0.5 mm. |
Cauline leaves | (8–)12–31(–43); sessile; blade ovate to lanceolate or oblong, margins usually dentate, rarely subentire, surfaces pubescent as basal. |
(0 or) 1 or 2 (3 or 4); sessile; blade elliptic to lanceolate or ovate, margins toothed, surfaces pubescent, abaxially with short-stalked 3- or 4-rayed trichomes, adaxially with simple and short-stalked, 2–4-rayed trichomes. |
Racemes | 10–52(–83)-flowered, usually ebracteate, rarely proximalmost 1 or 2 flowers bracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
(2–)4–10(–15)-flowered, ebracteate, elongated in fruit; rachis not flexuous, usually glabrous, rarely sparsely pubescent. |
Flowers | sepals oblong, 2.5–4 mm, pubescent, (trichomes simple and stalked, 2-rayed); petals yellow, oblanceolate, 5–7 × 1.5–2.2 mm; anthers oblong, 0.5–0.8 mm. |
(chasmogamous, petaliferous); sepals (green or purplish), oblong, 1.5–2.5 mm, glabrous or pubescent, (trichomes simple); petals yellow, oblanceolate, 2.5–3.5 × 0.6–1 mm; anthers ovate, 0.25–0.4 mm. |
Fruiting pedicels | horizontal to divaricate-ascending, straight or slightly curved upward, 4–10(–13) mm, pubescent as rachis abaxially, glabrous adaxially. |
horizontal to divaricate-ascending, straight, 3–14(–19) mm (subequaling or shorter than fruit), usually glabrous, rarely pubescent, trichomes 2–4-rayed. |
Fruits | (not appressed to rachis), lanceolate to ovate or oblong-lanceolate, slightly to strongly twisted or plane, flattened, 5–15 × 2–3.5 mm; valves puberulent, trichomes simple and subsessile, 2(–4)-rayed, 0.03–0.25(–0.8) mm; ovules 14–28 per ovary; style (1–)1.5–3.5 mm. |
linear or, rarely, linear-elliptic, plane, flattened, (8–)10–17(–20) × 1.5–2 mm; valves usually glabrous, rarely sparsely puberulent, trichomes simple; ovules 24–36 per ovary; style 0.01–0.15 mm. |
Seeds | oblong, 1–1.3 × 0.6–0.8 mm. |
ovoid, 0.9–1 × 0.5–0.6 mm. |
2n | = 18. |
= 40. |
Draba helleriana |
Draba stenoloba |
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Phenology | Flowering Jun–Sep. | Flowering Jun–Jul. |
Habitat | Oak and pine-fir woodlands, aspen groves, rocky meadows | Grassy knolls, glacial moraines and creek banks, mesic meadows, alpine thickets |
Elevation | 2100-3600 m (6900-11800 ft) | 600-2300 m (2000-7500 ft) |
Distribution |
AZ; CO; NM; Mexico (Nuevo León)
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AK; OR; WA; AB; BC; YT
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Discussion | Draba helleriana is highly variable and was divided by C. L. Hitchcock (1941) into four varieties. For a discussion of those and the circumscription of the species, see I. A. Al-Shehbaz and M. D. Windham (2007). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba stenoloba is occasionally confused with D. albertina, but is easily recognized by having exclusively 2–4-rayed (versus mostly simple) trichomes on stems proximally. It is rarely encountered and apparently confined to the Pacific Northwest. In contrast, D. albertina is common and widespread in the mountains of western North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 311. | FNA vol. 7, p. 340. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. aurea var. tylosa, D. helleriana var. leiocarpa, D. helleriana var. neomexicana, D. helleriana var. patens, D. helleriana var. pinetorum, D. neomexicana, D. neomexicana var. robusta, D. pallida, D. patens, D. pinetorum, D. stylosa | D. acinacis, D. hirta var. siliquosa, D. macouniana, D. nemorosa var. stenoloba, D. nitida var. praelonga, D. oligantha, D. stenoloba var. oligantha |
Name authority | Greene: Pittonia 4: 17. (1899) | Ledebour: Fl. Ross. 1: 154. (1841) |
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