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Heller's Draba

Carolina Draba, Carolina whitlow-grass, Carolina whitlow-mustard, creeping Draba

Habit Biennials or perennials; (short-lived); caudex simple or branched; not scapose. Annuals; scapose or subscapose.
Stems

branched, (0.1–)1.5–4.3(–5.1) dm, hirsute throughout, trichomes simple, 0.4–1.3(–1.8) mm, with short-stalked to subsessile, 3–5-rayed ones, 0.2–0.4 mm, (some slightly coarser, 2-rayed).

(simple or few from, or distal to, base), unbranched, (0.1–)0.3–1.2(–1.6) dm, sparsely to densely pubescent proximally, trichomes 2 (or 3)-rayed, 0.1–0.6 mm, sometimes with simple or spurred ones, to 0.9 mm, distally usually glabrous, rarely with few trichomes.

Basal leaves

rosulate;

petiolate;

petiole base ciliate or not;

blade oblanceolate to obovate, 0.9–4.1(–5.2) cm × 2–7(–10) mm, margins entire or dentate, surfaces pubescent, abaxially with stalked, cruciform, and fewer 3–5-rayed trichomes, 0.1–0.6 mm, adaxially with cruciform and simple trichomes, 0.4–1.3 mm, and smaller 2-rayed ones.

not rosulate;

petiole obscure;

blade elliptic or spatulate to obovate or suborbicular, 0.5–2.3(–3) cm × 1.5–8(–13) mm, margins entire, surfaces pubescent abaxially with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, adaxially with simple trichomes, 0.6–1 mm, with stalked, 2-rayed ones, to 0.7 mm.

Cauline leaves

(8–)12–31(–43);

sessile;

blade ovate to lanceolate or oblong, margins usually dentate, rarely subentire, surfaces pubescent as basal.

usually 0, rarely 1–3;

sessile;

blade similar to basal.

Racemes

10–52(–83)-flowered, usually ebracteate, rarely proximalmost 1 or 2 flowers bracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

(3–)5–12(–16)-flowered, ebracteate, (subumbellate), not elongated in fruit;

rachis not flexuous, usually glabrous, rarely sparsely pubescent.

Flowers

sepals oblong, 2.5–4 mm, pubescent, (trichomes simple and stalked, 2-rayed);

petals yellow, oblanceolate, 5–7 × 1.5–2.2 mm;

anthers oblong, 0.5–0.8 mm.

(late-season ones cleistogamous, apetalous);

sepals oblong, 1.5–2.3 mm, pubescent, (trichomes simple);

petals (rarely absent), white, spatulate, 2–4.5 × 1–1.5 mm;

anthers ovate, 0.4–0.5 mm.

Fruiting pedicels

horizontal to divaricate-ascending, straight or slightly curved upward, 4–10(–13) mm, pubescent as rachis abaxially, glabrous adaxially.

horizontal to divaricate-ascending, straight, (1–)2–7(–9) mm, glabrous or glabrate.

Fruits

(not appressed to rachis), lanceolate to ovate or oblong-lanceolate, slightly to strongly twisted or plane, flattened, 5–15 × 2–3.5 mm;

valves puberulent, trichomes simple and subsessile, 2(–4)-rayed, 0.03–0.25(–0.8) mm;

ovules 14–28 per ovary;

style (1–)1.5–3.5 mm.

linear to linear-oblong, plane, flattened, (5–)7–16(–20) × 1.2–2.3 mm;

valves glabrous or pubescent, trichomes usually simple and antrorse, 0.1–0.3 mm, rarely with fewer, spurred or 2-rayed ones;

ovules 32–88 per ovary;

style 0.02–0.1 mm.

Seeds

oblong, 1–1.3 × 0.6–0.8 mm.

oblong to ovoid, 0.5–0.8 × 0.3–0.5 mm.

2n

= 18.

= 16, 30, 32.

Draba helleriana

Draba reptans

Phenology Flowering Jun–Sep. Flowering Feb–Jun(-Aug).
Habitat Oak and pine-fir woodlands, aspen groves, rocky meadows Rock outcrops, dry slopes and hillsides, prairies, glades, roadsides, disturbed sites
Elevation 2100-3600 m (6900-11800 ft) 0-3000 m (0-9800 ft)
Distribution
from FNA
AZ; CO; NM; Mexico (Nuevo León)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; GA; IA; ID; IL; KS; MA; MI; MN; MO; MT; NC; ND; NE; NM; NY; OH; OK; OR; PA; SC; SD; TN; TX; UT; WA; WI; WY; AB; BC; MB; ON; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Draba helleriana is highly variable and was divided by C. L. Hitchcock (1941) into four varieties. For a discussion of those and the circumscription of the species, see I. A. Al-Shehbaz and M. D. Windham (2007).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba reptans is often confused with D. cuneifolia, but the two are easily separated. The rachises and pedicels of D. reptans are usually glabrous (rarely with a few isolated trichomes); those of D. cuneifolia are always densely pubescent. Interestingly, both species show parallel variations in chromosome number; it is currently unclear whether this variation is real or the result of misidentified specimens and/or erroneous counts.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 311. FNA vol. 7, p. 333.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. aurea var. tylosa, D. helleriana var. leiocarpa, D. helleriana var. neomexicana, D. helleriana var. patens, D. helleriana var. pinetorum, D. neomexicana, D. neomexicana var. robusta, D. pallida, D. patens, D. pinetorum, D. stylosa Arabis reptans, D. caroliniana, D. caroliniana var. dolichocarpa, D. caroliniana var. hunteri, D. caroliniana var. micrantha, D. caroliniana subsp. stellifera, D. caroliniana var. umbellata, D. coloradensis, D. hispidula, D. micrantha, D. reptans var. micrantha, D. reptans subsp. stellifera, D. reptans var. stellifera, D. umbellata, Tomostima caroliniana, Tomostima hispidula, Tomostima micrantha
Name authority Greene: Pittonia 4: 17. (1899) (Lamarck) Fernald: Rhodora 36: 368. (1934)
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