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Heller's Draba
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Porsild's Draba, Porsild's whitlow-grass
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| Biennials or perennials; (short-lived); caudex simple or branched; not scapose. |
Perennials; (cespitose); caudex branched (with persistent leaf remains); often scapose. |
branched, (0.1–)1.5–4.3(–5.1) dm, hirsute throughout, trichomes simple, 0.4–1.3(–1.8) mm, with short-stalked to subsessile, 3–5-rayed ones, 0.2–0.4 mm, (some slightly coarser, 2-rayed). |
unbranched, 0.2–1.2 dm, sparsely pubescent proximally, trichomes short-stalked, 3–5- rayed, (non-crisped), 0.2–0.4 mm, glabrous distally. |
rosulate; petiolate; petiole base ciliate or not; blade oblanceolate to obovate, 0.9–4.1(–5.2) cm × 2–7(–10) mm, margins entire or dentate, surfaces pubescent, abaxially with stalked, cruciform, and fewer 3–5-rayed trichomes, 0.1–0.6 mm, adaxially with cruciform and simple trichomes, 0.4–1.3 mm, and smaller 2-rayed ones. |
rosulate; subsessile or shortly petiolate; petiole base and margin ciliate proximally, (trichomes simple, subsetiform, stiff, 0.3–0.9 mm); blade oblanceolate to obovate, 0.4–1.5 cm × 1–3 mm, margins entire, surfaces pubescent, abaxially with stalked, 6–12-rayed, (non-crisped) trichomes, 0.2–0.5 mm, (midvein obscure), adaxially similar, or with simple trichomes on proximal 1/2. |
(8–)12–31(–43); sessile; blade ovate to lanceolate or oblong, margins usually dentate, rarely subentire, surfaces pubescent as basal. |
0 or 1; sessile; blade oblong to lanceolate, margins entire, surfaces pubescent as basal. |
10–52(–83)-flowered, usually ebracteate, rarely proximalmost 1 or 2 flowers bracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
3–13-flowered, ebracteate, elongated in fruit; rachis not flexuous, glabrous. |
sepals oblong, 2.5–4 mm, pubescent, (trichomes simple and stalked, 2-rayed); petals yellow, oblanceolate, 5–7 × 1.5–2.2 mm; anthers oblong, 0.5–0.8 mm. |
sepals ovate, 1.5–2 mm, pubescent, (trichomes simple and 2-rayed); petals white, spatulate, 2–4 × 1–1.7 mm; anthers ovate, ca. 0.2 mm. |
horizontal to divaricate-ascending, straight or slightly curved upward, 4–10(–13) mm, pubescent as rachis abaxially, glabrous adaxially. |
divaricate-ascending, straight, 1–4(–8) mm, glabrous. |
(not appressed to rachis), lanceolate to ovate or oblong-lanceolate, slightly to strongly twisted or plane, flattened, 5–15 × 2–3.5 mm; valves puberulent, trichomes simple and subsessile, 2(–4)-rayed, 0.03–0.25(–0.8) mm; ovules 14–28 per ovary; style (1–)1.5–3.5 mm. |
ovate to oblong, plane, flattened, 4–7.5 × 1.7–3 mm; valves glabrous; ovules 12–16 per ovary; style 0.1–0.3(–0.5) mm. |
oblong, 1–1.3 × 0.6–0.8 mm. |
ovoid, 0.7–1 × 0.5–0.6 mm. |
= 18. |
= 32. |
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| Flowering Jun–Sep. |
Flowering Jun–Jul. |
| Oak and pine-fir woodlands, aspen groves, rocky meadows |
Rock outcrops, talus, meadows, gravel slopes |
| 2100-3600 m (6900-11800 ft) |
600-3000 m (2000-9800 ft) |
|
AZ; CO; NM; Mexico (Nuevo León)
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AK; MT; WY; AB; YT |
Draba helleriana is highly variable and was divided by C. L. Hitchcock (1941) into four varieties. For a discussion of those and the circumscription of the species, see I. A. Al-Shehbaz and M. D. Windham (2007). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
According to H. H. Grundt et al. (2004), tetraploid Draba porsildii probably originated from a diploid ancestor such as D. lonchocarpa. The Wyoming plants are a close match to the type of the species, though they have distinctly shorter pedicels. Some of the Alaskan plants are more problematic; they have finer and more branched trichomes, larger flowers, longer pedicels, and longer styles than the type. It is likely that they belong to another species; detailed molecular and cytological studies on this complex are needed before any meaningful conclusions are reached. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 7, p. 311. |
FNA vol. 7, p. 331. |
| Brassicaceae > tribe Arabideae > Draba |
Brassicaceae > tribe Arabideae > Draba |
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis |
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis |
| D. aurea var. tylosa, D. helleriana var. leiocarpa, D. helleriana var. neomexicana, D. helleriana var. patens, D. helleriana var. pinetorum, D. neomexicana, D. neomexicana var. robusta, D. pallida, D. patens, D. pinetorum, D. stylosa |
D. nivalis var. brevicula, D. porsildii var. brevicula |
| Greene: Pittonia 4: 17. (1899) |
G. A. Mulligan: Canad. J. Bot. 52: 1795, fig. 8. (1974) |
| |
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