Draba helleriana |
Draba californica |
|
---|---|---|
Heller's Draba |
California Draba |
|
Habit | Biennials or perennials; (short-lived); caudex simple or branched; not scapose. | Perennials; (short-lived); caudex simple or branched; sometimes scapose. |
Stems | branched, (0.1–)1.5–4.3(–5.1) dm, hirsute throughout, trichomes simple, 0.4–1.3(–1.8) mm, with short-stalked to subsessile, 3–5-rayed ones, 0.2–0.4 mm, (some slightly coarser, 2-rayed). |
unbranched or branched (few), (0.2–)0.4–0.9(–1.2) dm, pubescent throughout, trichomes stellate, 4–8-rayed, 0.05–0.5 mm. |
Basal leaves | rosulate; petiolate; petiole base ciliate or not; blade oblanceolate to obovate, 0.9–4.1(–5.2) cm × 2–7(–10) mm, margins entire or dentate, surfaces pubescent, abaxially with stalked, cruciform, and fewer 3–5-rayed trichomes, 0.1–0.6 mm, adaxially with cruciform and simple trichomes, 0.4–1.3 mm, and smaller 2-rayed ones. |
rosulate; shortly petiolate; petiole ciliate, margin not ciliate, (trichomes simple and 2-rayed, 0.3–1 mm); blade oblanceolate, 0.6–2 cm × 1.5–4(–6) mm, margins entire, surfaces pubescent with short-stalked, 4–8-rayed trichomes, 0.1–0.4 mm. |
Cauline leaves | (8–)12–31(–43); sessile; blade ovate to lanceolate or oblong, margins usually dentate, rarely subentire, surfaces pubescent as basal. |
0–3; sessile; blade ovate to oblong, margins entire, surfaces pubescent as basal. |
Racemes | 10–52(–83)-flowered, usually ebracteate, rarely proximalmost 1 or 2 flowers bracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
3–13-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals oblong, 2.5–4 mm, pubescent, (trichomes simple and stalked, 2-rayed); petals yellow, oblanceolate, 5–7 × 1.5–2.2 mm; anthers oblong, 0.5–0.8 mm. |
sepals (persistent until fruit maturity), oblong to ovate, 1.7–2.5 mm, pubescent, (trichomes short-stalked, 2–5-rayed); petals white or creamy white, spatulate to oblanceolate, 2.2–3 × 0.8–1 mm; anthers ovate, 0.2–0.35 mm. |
Fruiting pedicels | horizontal to divaricate-ascending, straight or slightly curved upward, 4–10(–13) mm, pubescent as rachis abaxially, glabrous adaxially. |
ascending to divaricate-ascending, straight, (2–)4–9(–12) mm, pubescent, trichomes short-stalked, 4–8-rayed, (0.1–0.4 mm). |
Fruits | (not appressed to rachis), lanceolate to ovate or oblong-lanceolate, slightly to strongly twisted or plane, flattened, 5–15 × 2–3.5 mm; valves puberulent, trichomes simple and subsessile, 2(–4)-rayed, 0.03–0.25(–0.8) mm; ovules 14–28 per ovary; style (1–)1.5–3.5 mm. |
lanceolate to oblong, plane, slightly flattened, (5–)6–9(–11) × 1.8–2.5 mm; valves usually pubescent, rarely glabrous, trichomes simple and short-stalked, 2- or 3-rayed, 0.05–0.25 mm; ovules 22–32 per ovary; style 0.1–0.3(–0.4) mm. |
Seeds | oblong, 1–1.3 × 0.6–0.8 mm. |
oblong, 0.8–1 × 0.5–0.6 mm. |
2n | = 18. |
|
Draba helleriana |
Draba californica |
|
Phenology | Flowering Jun–Sep. | Flowering Jun–Aug. |
Habitat | Oak and pine-fir woodlands, aspen groves, rocky meadows | Grassy meadows, alpine areas, fellfields |
Elevation | 2100-3600 m (6900-11800 ft) | 3200-4000 m (10500-13100 ft) |
Distribution |
AZ; CO; NM; Mexico (Nuevo León)
|
CA; NV |
Discussion | Draba helleriana is highly variable and was divided by C. L. Hitchcock (1941) into four varieties. For a discussion of those and the circumscription of the species, see I. A. Al-Shehbaz and M. D. Windham (2007). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although originally treated as a variety of Draba cuneifolia, D. californica is but distantly related to that species. Instead, ongoing studies (L. Allphin and M. D. Windham, unpubl.) suggest that it may be an allopolyploid resulting from hybridization between D. albertina and D. breweri. The species is restricted to the White Mountains in Mono County, California, and adjacent Esmeralda County, Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 311. | FNA vol. 7, p. 298. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. aurea var. tylosa, D. helleriana var. leiocarpa, D. helleriana var. neomexicana, D. helleriana var. patens, D. helleriana var. pinetorum, D. neomexicana, D. neomexicana var. robusta, D. pallida, D. patens, D. pinetorum, D. stylosa | D. cuneifolia var. californica |
Name authority | Greene: Pittonia 4: 17. (1899) | (Jepson) Rollins & R. A. Price: Aliso 12: 19. (1988) |
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