Draba helleriana |
Draba asprella |
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Heller's Draba |
rough Draba, rough whitlowgrass |
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Habit | Biennials or perennials; (short-lived); caudex simple or branched; not scapose. | Perennials; (cespitose); caudex branched (with some persistent leaf bases); scapose. | ||||
Stems | branched, (0.1–)1.5–4.3(–5.1) dm, hirsute throughout, trichomes simple, 0.4–1.3(–1.8) mm, with short-stalked to subsessile, 3–5-rayed ones, 0.2–0.4 mm, (some slightly coarser, 2-rayed). |
unbranched, (0.4–)0.6–1.5(–2.7) dm, pubescent throughout, trichomes simple, 0.5–1.5 mm, and 2–4-rayed ones, 0.05–0.3 mm, (sometimes 2–4-rayed ones distally). |
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Basal leaves | rosulate; petiolate; petiole base ciliate or not; blade oblanceolate to obovate, 0.9–4.1(–5.2) cm × 2–7(–10) mm, margins entire or dentate, surfaces pubescent, abaxially with stalked, cruciform, and fewer 3–5-rayed trichomes, 0.1–0.6 mm, adaxially with cruciform and simple trichomes, 0.4–1.3 mm, and smaller 2-rayed ones. |
rosulate; subsessile or shortly petiolate; petiole ciliate throughout; blade oblanceolate to spatulate or obovate, (0.6–)0.8–4.5(–6) cm × 2–12(–16) mm, margins usually entire, rarely obscurely dentate, (not ciliate), surfaces abaxially pubescent with stalked, (2–)4-rayed trichomes, 0.2–1 mm, adaxially with stalked, 2–4-rayed trichomes, 0.3–0.8 mm, and simple ones, 0.5–1.9 mm. |
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Cauline leaves | (8–)12–31(–43); sessile; blade ovate to lanceolate or oblong, margins usually dentate, rarely subentire, surfaces pubescent as basal. |
0. |
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Racemes | 10–52(–83)-flowered, usually ebracteate, rarely proximalmost 1 or 2 flowers bracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
(15–)30–75-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent, trichomes 2–4-rayed, sometimes with simple ones. |
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Flowers | sepals oblong, 2.5–4 mm, pubescent, (trichomes simple and stalked, 2-rayed); petals yellow, oblanceolate, 5–7 × 1.5–2.2 mm; anthers oblong, 0.5–0.8 mm. |
sepals broadly ovate, 1.8–3.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals pale yellow, oblanceolate, 4–7 × 1.5–2.5 mm; anthers ovate to oblong, 0.4–0.7 mm. |
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Fruiting pedicels | horizontal to divaricate-ascending, straight or slightly curved upward, 4–10(–13) mm, pubescent as rachis abaxially, glabrous adaxially. |
horizontal to divaricate-ascending, straight, 3–11(–16) mm, pubescent, trichomes 2–4-rayed, sometimes with simple ones. |
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Fruits | (not appressed to rachis), lanceolate to ovate or oblong-lanceolate, slightly to strongly twisted or plane, flattened, 5–15 × 2–3.5 mm; valves puberulent, trichomes simple and subsessile, 2(–4)-rayed, 0.03–0.25(–0.8) mm; ovules 14–28 per ovary; style (1–)1.5–3.5 mm. |
ovoid-ellipsoid to oblong, plane, inflated basally, 3–8 × 1.8–3.5 mm; valves hirsute, trichomes simple and spurred, (0.2–)0.3–1 mm, or puberulent, trichomes short-stalked, 2–4-rayed, 0.1–0.2 mm; ovules 8–12(–18) per ovary; style 0.6–2(–2.5) mm. |
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Seeds | oblong, 1–1.3 × 0.6–0.8 mm. |
oblong, 0.9–1.2 × 0.6–0.8 mm. |
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2n | = 18. |
= 30. |
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Draba helleriana |
Draba asprella |
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Phenology | Flowering Jun–Sep. | |||||
Habitat | Oak and pine-fir woodlands, aspen groves, rocky meadows | |||||
Elevation | 2100-3600 m (6900-11800 ft) | |||||
Distribution |
AZ; CO; NM; Mexico (Nuevo León)
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AZ
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Discussion | Draba helleriana is highly variable and was divided by C. L. Hitchcock (1941) into four varieties. For a discussion of those and the circumscription of the species, see I. A. Al-Shehbaz and M. D. Windham (2007). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Both R. C. Rollins (1993) and N. H. Holmgren (2005b) indicated that the chromosome number of Draba asprella is 2n = 32, but repeated counts by one of us (M. D. Windham 2000, unpubl.) show that the species consistently has 2n = 30. Rollins divided D. asprella into four varieties encompassing tremendous variation in trichome morphology. One of these (var. zionensis) is more closely related to D. sobolifera (Windham and L. Allphin, unpubl.) and is treated herein at species rank. Within D. asprella, in the strict sense, we recognize two varieties separated primarily by the type of trichomes found on the fruits. The distinctions are not absolute and there appear to be forms connecting the two, especially in Coconino County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 311. | FNA vol. 7, p. 293. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | D. aurea var. tylosa, D. helleriana var. leiocarpa, D. helleriana var. neomexicana, D. helleriana var. patens, D. helleriana var. pinetorum, D. neomexicana, D. neomexicana var. robusta, D. pallida, D. patens, D. pinetorum, D. stylosa | |||||
Name authority | Greene: Pittonia 4: 17. (1899) | Greene: Bull. Torrey Bot. Club 10: 125. (1883) | ||||
Web links |