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Heller's Draba
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desert Draba
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| Biennials or perennials; (short-lived); caudex simple or branched; not scapose. |
Perennials; (cespitose); caudex branched (with persistent leaf remains, branches relatively short); not scapose. |
branched, (0.1–)1.5–4.3(–5.1) dm, hirsute throughout, trichomes simple, 0.4–1.3(–1.8) mm, with short-stalked to subsessile, 3–5-rayed ones, 0.2–0.4 mm, (some slightly coarser, 2-rayed). |
usually unbranched, (0.3–)0.4–1 dm, pubescent throughout, trichomes simple and 2-rayed, 0.4–1.4 mm, and shorter, 2–4-rayed ones, 0.05–0.3 mm. |
rosulate; petiolate; petiole base ciliate or not; blade oblanceolate to obovate, 0.9–4.1(–5.2) cm × 2–7(–10) mm, margins entire or dentate, surfaces pubescent, abaxially with stalked, cruciform, and fewer 3–5-rayed trichomes, 0.1–0.6 mm, adaxially with cruciform and simple trichomes, 0.4–1.3 mm, and smaller 2-rayed ones. |
rosulate; subsessile; petiole not ciliate; blade obovate to spatulate or oblanceolate, 0.6–1.9(–2.4) cm × 2–5(–8) mm, (base ciliate, trichomes simple and 2-rayed), margins entire, surfaces abaxially pubescent with stalked, (2–)4-rayed trichomes, 0.3–0.6 mm, adaxially with simple and stalked, 2-rayed trichomes, 0.3–0.6 mm, and smaller, 3- or 4-rayed ones. |
(8–)12–31(–43); sessile; blade ovate to lanceolate or oblong, margins usually dentate, rarely subentire, surfaces pubescent as basal. |
(1 or) 2–5; sessile; blade oblong to oblanceolate or ovate, margins entire, surfaces pubescent as basal. |
10–52(–83)-flowered, usually ebracteate, rarely proximalmost 1 or 2 flowers bracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
12–40-flowered, ebracteate, slightly elongated in fruit; rachis not flexuous, pubescent, trichomes simple and 2–4-rayed. |
sepals oblong, 2.5–4 mm, pubescent, (trichomes simple and stalked, 2-rayed); petals yellow, oblanceolate, 5–7 × 1.5–2.2 mm; anthers oblong, 0.5–0.8 mm. |
sepals broadly ovate, 2–2.7 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals yellow, spatulate to oblanceolate, 3.5–5 × 1.4–2 mm; anthers oblong, 0.4–0.6 mm. |
horizontal to divaricate-ascending, straight or slightly curved upward, 4–10(–13) mm, pubescent as rachis abaxially, glabrous adaxially. |
horizontal to divaricate-ascending, straight, (3–)4–8 mm, pubescent, trichomes simple and 2–4-rayed (0.1–0.5 mm). |
(not appressed to rachis), lanceolate to ovate or oblong-lanceolate, slightly to strongly twisted or plane, flattened, 5–15 × 2–3.5 mm; valves puberulent, trichomes simple and subsessile, 2(–4)-rayed, 0.03–0.25(–0.8) mm; ovules 14–28 per ovary; style (1–)1.5–3.5 mm. |
elliptic or ovate to lanceolate, plane, slightly inflated basally, 4–7 × 2.5–3.5 mm; valves pubescent, trichomes simple and stalked, 2–4-rayed, 0.05–0.4 mm; ovules 8–12 per ovary; style 0.5–1.2(–1.4) mm. |
oblong, 1–1.3 × 0.6–0.8 mm. |
ovoid, 1–1.4 × 0.6–0.8 mm. |
= 18. |
= 24. |
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| Flowering Jun–Sep. |
Flowering May–Jul. |
| Oak and pine-fir woodlands, aspen groves, rocky meadows |
Rock crevices and gravelly soil in conifer and subalpine shrub communities |
| 2100-3600 m (6900-11800 ft) |
2100-3400 m (6900-11200 ft) |
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AZ; CO; NM; Mexico (Nuevo León)
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NV |
Draba helleriana is highly variable and was divided by C. L. Hitchcock (1941) into four varieties. For a discussion of those and the circumscription of the species, see I. A. Al-Shehbaz and M. D. Windham (2007). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Draba arida is a sexually reproducing, diploid member of the D. ventosa complex that may have been involved in the origin of the more widespread apomictic triploids (M. A. Beilstein and M. D. Windham 2003). The species is known from the Monitor, Toiyabe, and Toquima ranges in Lander and Nye counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 7, p. 311. |
FNA vol. 7, p. 292. |
| Brassicaceae > tribe Arabideae > Draba |
Brassicaceae > tribe Arabideae > Draba |
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis |
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis |
| D. aurea var. tylosa, D. helleriana var. leiocarpa, D. helleriana var. neomexicana, D. helleriana var. patens, D. helleriana var. pinetorum, D. neomexicana, D. neomexicana var. robusta, D. pallida, D. patens, D. pinetorum, D. stylosa |
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| Greene: Pittonia 4: 17. (1899) |
C. L. Hitchcock: Revis. Drabas W. N. Amer., 52, plate 3, fig. 23. (1941) |
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