Draba heilii |
Draba cuneifolia |
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wedge leaf Draba |
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Habit | Perennials; (densely cespitose); caudex few-branched (compact, with persistent, thickened leaves); not scapose. | Annuals; scapose or subscapose. | ||||||||
Stems | unbranched or branched distally, 0.3–0.7 dm, glabrous throughout or, rarely, pilose, trichomes simple and stalked, 2-rayed, 0.2–0.5 mm. |
(simple to many from or near base), unbranched, (0.2–) 0.3–2.7(–3.7) dm, hirsute or pubescent throughout, trichomes 3–4-rayed, 0.05–0.4 mm, (sometimes mixed proximally with simple or spurred ones, 0.5–1.2 mm). |
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Basal leaves | (persistent, imbricate); rosulate; undifferentiated into blade and petiole (becoming indurate at base); blade linear-lanceolate, 0.5–0.8 cm × 1–1.5(–2) mm, margins entire, surfaces glabrous or, rarely, pilose with simple and stalked, 2-rayed trichomes, (margins ciliate with simple and, rarely, 2-rayed setiform trichomes, 0.5–1.3 mm, midvein prominent). |
not rosulate; petiole (obscure), not ciliate; blade oblanceolate to spatulate or broadly obovate, (0.4–)1–3.5(–5) cm × (2–)6–20(–28) mm, margins dentate (in distal 1/2), surfaces pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.1–0.7 mm, adaxially similar or also with fewer, simple trichomes, 0.4–0.7 mm. |
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Cauline leaves | 6–8; sessile; blade linear-oblong, 0.5–0.8 cm × 1–2 mm (widest about base), margins entire, (ciliate). |
0–6 (on proximal 1/3 of stem); blade similar to basal. |
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Racemes | 10–26-flowered, ebracteate, elongated in fruit; rachis not flexuous, usually glabrous, rarely pubescent, (trichomes non-crisped, simple and 2-rayed). |
10–50(–70)-flowered (throughout or on distal 1/3 of scape), ebracteate, elongated in fruit; rachis not flexuous, densely pubescent, trichomes 2–4-rayed. |
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Flowers | sepals ovate, 1.7–2.7 mm, usually glabrous, rarely pilose, (trichomes simple); petals yellow, oblong-oblanceolate, 4–6 × 1.5–2 mm; anthers oblong, 0.5–0.6 mm. |
(late ones cleistogamous, apetalous); sepals (green or pink), oblong, 1.5–2.5 mm, glabrous or pubescent, (trichomes simple); petals white, spatulate, (2–)2.5–4.5(–5) × 1–2 mm, (emarginate or obtuse); anthers ovate to oblong, (0.1–)0.25–0.4 mm. |
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Fruiting pedicels | ascending, straight, 3–5(–7) mm, usually glabrous, rarely pilose, (trichomes simple and 2-rayed, stalked). |
horizontal to divaricate-ascending, straight, (1–)2–7(–10) mm, pubescent as rachis. |
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Fruits | narrowly lanceolate, twisted 1/2 turn or plane, flattened, 7–10 × 1.3–1.6 mm; valves glabrous; ovules 16–20 per ovary; style 0.7–1.3 mm. |
oblong to linear or lanceolate to broadly ovate, plane, flattened, (3–)6–12(–16) × 1.7–2.7(–3) mm; valves usually puberulent, rarely glabrous, trichomes simple, antrorse, 0.1–0.3 mm, (rarely with 2-rayed ones, or all trichomes short-stalked, 4-rayed, cruciform); ovules (12–)24–66(–72) per ovary; style 0.01–0.3(–0.4) mm. |
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Seeds | ovoid, 1 × 0.6 mm. |
broadly ovoid, 0.5–0.7 × 0.4–0.5 mm. |
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2n | = 30, 32. |
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Draba heilii |
Draba cuneifolia |
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Phenology | Flowering Jun–Jul. | |||||||||
Habitat | Alpine tundra | |||||||||
Elevation | ca. 3700 m (ca. 12100 ft) | |||||||||
Distribution |
NM |
AL; AR; AZ; CA; CO; FL; IL; KS; KY; LA; MO; MS; NC; NE; NM; NV; OH; OK; PA; SD; TN; TX; UT; n Mexico |
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Discussion | Draba heilii is restricted to Mora and Rio Arriba counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). Draba cuneifolia is highly variable in fruit shape and size, style length, ovule number per ovary, type of fruit indumentum, and length of the fruiting raceme in relation to the rest of the scape. Reported chromosome numbers also vary, but it is currently unclear whether this is the result of biologically relevant processes or erroneous counts. R. L. Hartman et al. (1975) divided the species into three varieties, a position followed by subsequent authors (e.g., R. C. Rollins 1993). Typical var. sonorae is rather distinct (see key below) and may deserve recognition as a separate species, as by O. E. Schulz (1927). Critical, population-based, molecular studies are needed to assess this. In the absence of such studies, we tentatively follow Hartman et al., recognizing three varieties that are distinguished primarily by the types of trichomes found on the fruits. Glabrous-fruited forms occasionally encountered in all three varieties can be difficult to identify. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 311. | FNA vol. 7, p. 303. | ||||||||
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Name authority | Al-Shehbaz: Harvard Pap. Bot. 14: 83, fig. 1. (2009) | Nuttall ex Torrey & A. Gray: Fl. N. Amer. 1: 108. (1838) | ||||||||
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