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wedge leaf Draba

Habit Perennials; (densely cespitose); caudex few-branched (compact, with persistent, thickened leaves); not scapose. Annuals; scapose or subscapose.
Stems

unbranched or branched distally, 0.3–0.7 dm, glabrous throughout or, rarely, pilose, trichomes simple and stalked, 2-rayed, 0.2–0.5 mm.

(simple to many from or near base), unbranched, (0.2–) 0.3–2.7(–3.7) dm, hirsute or pubescent throughout, trichomes 3–4-rayed, 0.05–0.4 mm, (sometimes mixed proximally with simple or spurred ones, 0.5–1.2 mm).

Basal leaves

(persistent, imbricate); rosulate; undifferentiated into blade and petiole (becoming indurate at base);

blade linear-lanceolate, 0.5–0.8 cm × 1–1.5(–2) mm, margins entire, surfaces glabrous or, rarely, pilose with simple and stalked, 2-rayed trichomes, (margins ciliate with simple and, rarely, 2-rayed setiform trichomes, 0.5–1.3 mm, midvein prominent).

not rosulate;

petiole (obscure), not ciliate;

blade oblanceolate to spatulate or broadly obovate, (0.4–)1–3.5(–5) cm × (2–)6–20(–28) mm, margins dentate (in distal 1/2), surfaces pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.1–0.7 mm, adaxially similar or also with fewer, simple trichomes, 0.4–0.7 mm.

Cauline leaves

6–8;

sessile;

blade linear-oblong, 0.5–0.8 cm × 1–2 mm (widest about base), margins entire, (ciliate).

0–6 (on proximal 1/3 of stem);

blade similar to basal.

Racemes

10–26-flowered, ebracteate, elongated in fruit;

rachis not flexuous, usually glabrous, rarely pubescent, (trichomes non-crisped, simple and 2-rayed).

10–50(–70)-flowered (throughout or on distal 1/3 of scape), ebracteate, elongated in fruit;

rachis not flexuous, densely pubescent, trichomes 2–4-rayed.

Flowers

sepals ovate, 1.7–2.7 mm, usually glabrous, rarely pilose, (trichomes simple);

petals yellow, oblong-oblanceolate, 4–6 × 1.5–2 mm;

anthers oblong, 0.5–0.6 mm.

(late ones cleistogamous, apetalous);

sepals (green or pink), oblong, 1.5–2.5 mm, glabrous or pubescent, (trichomes simple);

petals white, spatulate, (2–)2.5–4.5(–5) × 1–2 mm, (emarginate or obtuse);

anthers ovate to oblong, (0.1–)0.25–0.4 mm.

Fruiting pedicels

ascending, straight, 3–5(–7) mm, usually glabrous, rarely pilose, (trichomes simple and 2-rayed, stalked).

horizontal to divaricate-ascending, straight, (1–)2–7(–10) mm, pubescent as rachis.

Fruits

narrowly lanceolate, twisted 1/2 turn or plane, flattened, 7–10 × 1.3–1.6 mm;

valves glabrous;

ovules 16–20 per ovary;

style 0.7–1.3 mm.

oblong to linear or lanceolate to broadly ovate, plane, flattened, (3–)6–12(–16) × 1.7–2.7(–3) mm;

valves usually puberulent, rarely glabrous, trichomes simple, antrorse, 0.1–0.3 mm, (rarely with 2-rayed ones, or all trichomes short-stalked, 4-rayed, cruciform);

ovules (12–)24–66(–72) per ovary;

style 0.01–0.3(–0.4) mm.

Seeds

ovoid, 1 × 0.6 mm.

broadly ovoid, 0.5–0.7 × 0.4–0.5 mm.

2n

= 30, 32.

Draba heilii

Draba cuneifolia

Phenology Flowering Jun–Jul.
Habitat Alpine tundra
Elevation ca. 3700 m (ca. 12100 ft)
Distribution
from FNA
NM
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; FL; IL; KS; KY; LA; MO; MS; NC; NE; NM; NV; OH; OK; PA; SD; TN; TX; UT; n Mexico
[BONAP county map]
Discussion

Draba heilii is restricted to Mora and Rio Arriba counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Draba cuneifolia is highly variable in fruit shape and size, style length, ovule number per ovary, type of fruit indumentum, and length of the fruiting raceme in relation to the rest of the scape. Reported chromosome numbers also vary, but it is currently unclear whether this is the result of biologically relevant processes or erroneous counts. R. L. Hartman et al. (1975) divided the species into three varieties, a position followed by subsequent authors (e.g., R. C. Rollins 1993). Typical var. sonorae is rather distinct (see key below) and may deserve recognition as a separate species, as by O. E. Schulz (1927). Critical, population-based, molecular studies are needed to assess this. In the absence of such studies, we tentatively follow Hartman et al., recognizing three varieties that are distinguished primarily by the types of trichomes found on the fruits. Glabrous-fruited forms occasionally encountered in all three varieties can be difficult to identify.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit valves: trichomes usually simple, sometimes with 2-rayed ones; stems hirsute with at least some simple trichomes.
var. cuneifolia
1. Fruit valves: trichomes stalked and 4-rayed, sometimes with 2- or 3-rayed ones; stems pubescent with 3- or 4-rayed trichomes
→ 2
2. Fruits (5-)7-12 mm; ovules 20-44 per ovary; racemes to 3/4 of scape.
var. integrifolia
2. Fruits 3-6(-8) mm; ovules 12-24 per ovary; racemes on all or most of scape.
var. sonorae
Source FNA vol. 7, p. 311. FNA vol. 7, p. 303.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Subordinate taxa
D. cuneifolia var. cuneifolia, D. cuneifolia var. integrifolia, D. cuneifolia var. sonorae
Name authority Al-Shehbaz: Harvard Pap. Bot. 14: 83, fig. 1. (2009) Nuttall ex Torrey & A. Gray: Fl. N. Amer. 1: 108. (1838)
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