Draba heilii |
Draba alpina |
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alpine Draba |
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Habit | Perennials; (densely cespitose); caudex few-branched (compact, with persistent, thickened leaves); not scapose. | Perennials; (cespitose); caudex branched (covered with persistent leaves or leaf remains); scapose. |
Stems | unbranched or branched distally, 0.3–0.7 dm, glabrous throughout or, rarely, pilose, trichomes simple and stalked, 2-rayed, 0.2–0.5 mm. |
unbranched, (0.3–)0.5–1.7(–2.8) dm, pubescent throughout, trichomes simple and 2-rayed, 0.3–0.8 mm, with 3–5-rayed ones, 0.1–0.3 mm. |
Basal leaves | (persistent, imbricate); rosulate; undifferentiated into blade and petiole (becoming indurate at base); blade linear-lanceolate, 0.5–0.8 cm × 1–1.5(–2) mm, margins entire, surfaces glabrous or, rarely, pilose with simple and stalked, 2-rayed trichomes, (margins ciliate with simple and, rarely, 2-rayed setiform trichomes, 0.5–1.3 mm, midvein prominent). |
rosulate; petiolate; petiole base (not thickened), ciliate, (trichomes simple, 0.3–1 mm); blade oblanceolate to obovate or lanceolate to oblong, 0.8–3(–4.5) cm × 2.5–6(–9) mm, margins entire, surfaces abaxially pubescent with stalked, 2–4-rayed trichomes, 0.1–0.5 mm, with simple ones (midvein obscure, not thickened), adaxially glabrous or sparsely pubescent with simple and stalked, 2–4-rayed trichomes. |
Cauline leaves | 6–8; sessile; blade linear-oblong, 0.5–0.8 cm × 1–2 mm (widest about base), margins entire, (ciliate). |
0. |
Racemes | 10–26-flowered, ebracteate, elongated in fruit; rachis not flexuous, usually glabrous, rarely pubescent, (trichomes non-crisped, simple and 2-rayed). |
6–18-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals ovate, 1.7–2.7 mm, usually glabrous, rarely pilose, (trichomes simple); petals yellow, oblong-oblanceolate, 4–6 × 1.5–2 mm; anthers oblong, 0.5–0.6 mm. |
sepals (purplish tinged), narrowly ovate, 2.5–3 mm, pubescent, (trichomes simple and fewer, stalked, 2-rayed); petals bright yellow, narrowly obovate, 3.5–5 × 1.7–2.5 mm; anthers ovate, 0.3–0.4 mm. |
Fruiting pedicels | ascending, straight, 3–5(–7) mm, usually glabrous, rarely pilose, (trichomes simple and 2-rayed, stalked). |
ascending to divaricate-ascending, straight or, sometimes, slightly curved upwards, 4–14(–30) mm, pubescent, trichomes simple and 2–4-rayed. |
Fruits | narrowly lanceolate, twisted 1/2 turn or plane, flattened, 7–10 × 1.3–1.6 mm; valves glabrous; ovules 16–20 per ovary; style 0.7–1.3 mm. |
elliptic, plane, flattened, 6–10 × 2–3 mm; valves glabrous or glabrescent, trichomes simple, (not confined to replum); ovules 12–24 per ovary; style 0.2–0.3 mm (stigma about as wide as style). |
Seeds | ovoid, 1 × 0.6 mm. |
(pale brown), ovoid, 0.9–1.3 × 0.6–0.9 mm. |
2n | = 80. |
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Draba heilii |
Draba alpina |
|
Phenology | Flowering Jun–Jul. | Flowering Jun–Aug. |
Habitat | Alpine tundra | Moist tundra and ridges, sand and gravel flats or beaches |
Elevation | ca. 3700 m (ca. 12100 ft) | 0-1000 m (0-3300 ft) |
Distribution |
NM |
LB; MB; NU; ON; QC; Greenland; Europe (Finland, Norway, Russia, Sweden)
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Discussion | Draba heilii is restricted to Mora and Rio Arriba counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The synonymy above includes two North American names overlooked by C. L. Hitchcock (1941) and R. C. Rollins (1993). Draba alpina was broadly delimited by O. E. Schulz (1927) and included 17 varieties, some of which (e.g., corymbosa, oxycarpa, pilosa) are recognized herein as distinct species. The name D. alpina was so misapplied that it was used for any circumpolar or alpine, scapose, yellow-flowered, perennial Draba. Various chromosome numbers (e.g., 2n = 64, 80, 112, 120; S. I. Warwick and I. A. Al-Shehbaz 2006) have been reported for the species. As circumscribed here, it has the narrow distribution outlined above and includes plants with 2n = 80. Reports of the species from Alaska, Canadian Northwest Territories and Yukon, Siberia, eastern Asia, Russian Far East, and the Central Asian republics are either suspect or very unlikely. The entire D. alpina complex (including the above three species, D. glacialis Adams, D. macounii, etc.) is in need of critical molecular, cytological, and morphological study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 311. | FNA vol. 7, p. 290. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. alpina var. hydeana, D. alpina var. inflatisiliqua | |
Name authority | Al-Shehbaz: Harvard Pap. Bot. 14: 83, fig. 1. (2009) | Linnaeus: Sp. Pl. 2: 642. (1753) |
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