Draba grayana |
Draba corymbosa |
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Gray's Draba |
Baffin bay Draba, Baffin bay whitlow-grass, flat-top Draba |
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Habit | Perennials; (densely cespitose); caudex branched (covered with persistent, somewhat thickened, dry petioles); not scapose. | Perennials; (cespitose); caudex branched (with persistent leaves or leaf remains, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, 0.08–0.6 dm, densely to moderately pubescent throughout, trichomes simple and stalked, 2- or 3-rayed, 0.2–1 mm. |
unbranched, (0.05–)0.2–0.8(–1.5) dm, pubescent throughout, trichomes simple and 2-rayed, 0.4–1 mm, with 3–5-rayed ones, 0.05–0.4 mm, (sometimes trichomes mostly simple). |
Basal leaves | rosulate; petiolate; petiole ciliate, (trichomes simple, 0.2–1 mm); blade linear to linear-oblanceolate, (0.4–)0.6–1.5(–2) cm × 1–2 mm, margins entire, (pubescent as petiole), surfaces pubescent abaxially with simple and stalked, 2- or 3-rayed trichomes, 0.1–0.4 mm, adaxially glabrate or subapically sparsely pubescent with simple trichomes. |
rosulate; petiolate; petiole base and margin ciliate, (trichomes often course, simple, 0.4–1.3 mm); blade oblanceolate to obovate, 0.6–1.8 cm × 1.5–5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–6-rayed trichomes, 0.2–0.6 mm, adaxially with primarily simple and stalked, 2-rayed trichomes, to 1.1 mm, with 3–5-rayed ones, 0.2–0.4 mm. |
Cauline leaves | 1–4; sessile; blade linear to linear-oblanceolate, margins entire, surfaces pubescent as basal. |
0. |
Racemes | 5–12(–16)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
2–9(–12)-flowered, ebracteate, (corymbose), slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals ovate, 1.5–2 mm, pubescent, (trichomes simple and short-stalked, 2- or 3-rayed); petals bright yellow, spatulate, 3–4.5 × 1–2 mm, (flared, clawed); anthers ovate, 0.3–0.5 mm. |
sepals (grayish green), broadly ovate, 2.2–3 mm, pubescent, (trichomes simple, to 1 mm, sometimes with stalked, smaller, 2–4-rayed ones); petals (broadly patent), yellow, obovate, 4–6 × 3–5 mm; anthers ovate, 0.3–0.4 mm. |
Fruiting pedicels | divaricate-ascending, straight or slightly curved upward, 1.5–6 mm, pubescent as stem. |
divaricate-ascending, straight or slightly curved upward, 4–11(–16) mm, pubescent, trichomes simple and 2–4-rayed. |
Fruits | lanceolate to oblong-lanceolate, slightly twisted or plane, flattened, 4–12 × 1.5–3 mm; valves glabrous or pubescent, trichomes simple, 0.07–0.2 mm; ovules 16–20 per ovary; style 0.4–1.2 mm. |
oblong or ovate, plane, flattened, 6–12 × 3.5–5.5 mm; valves pubescent or puberulent, trichomes simple, 0.1–0.4 mm, (sometimes with short-stalked, 2- or 3-rayed ones); ovules 12–24 per ovary; style 0.6–1 mm (stigma distinctly wider than style). |
Seeds | ovoid, 1–1.4 × 0.7–1 mm. |
(brown), ovoid, 1–1.3 × 0.6–0.9 mm. |
2n | = 24 ± 3. |
= 128, 144. |
Draba grayana |
Draba corymbosa |
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Phenology | Flowering Jul–Aug. | Flowering Jun–Aug. |
Habitat | Alpine tundra, fellfields, gravelly wet meadows | Moist tundra, among calcareous or dolomitic rocks, gravel beaches, silt and clay terraces |
Elevation | 3500-4000 m (11500-13100 ft) | 0-1700 m (0-5600 ft) |
Distribution |
CO |
AK; BC; NT; NU; YT; Greenland; Europe (Norway, n Russia); e Asia (Russian Far East, Siberia) |
Discussion | Of conservation concern. Draba grayana is an apomictic species clearly related to D. exunguiculata, with which it is occasionally sympatric (see discussion of 36. D. exunguiculata for distinguishing features). It is known from Chaffe, Clear Creek, Larimer, Park, and Summit counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba corymbosa, which is 16-ploid or 18-ploid with x = 8, is an extremely variable species of polyphyletic, allopolyploid origin. Most individuals appear to have decaploid D. alpina (2n = 80) in their parentage, but the other genomes are provided by hexaploid (2n = 48) and octoploid (2n = 64) species (C. Brochmann et al. 1993). O. E. Schulz reduced D. corymbosa to a variety of D. alpina; R. C. Rollins (1993) treated it as a distinct species. Draba corymbosa is distinguished from D. alpina by having pubescent or puberulent (versus glabrous or glabrescent) fruits, corymbose (versus usually elongated) fruiting racemes, and abaxial leaf blade surfaces always lacking (versus usually with some) simple trichomes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 310. | FNA vol. 7, p. 301. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. streptocarpa var. grayana, D. alpicola | D. alpina var. bellii, D. alpina var. corymbosa, D. barbata, D. bellii, D. kjellmanii, D. macrocarpa, D. vestita |
Name authority | (Rydberg) C. L. Hitchcock: Revis. Drabas W. N. Amer., 29. (1941) | R. Brown ex de Candolle: Syst. Nat. 2: 343. (1821) |
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