Draba grayana |
Draba cana |
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Gray's Draba |
Brewer's Draba, canescent Draba, canescent whitlow-mustard, cushion Draba, hoary Draba, hoary whitlow-grass, lance-leaf Draba, lance-leaf whitlow-grass |
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Habit | Perennials; (densely cespitose); caudex branched (covered with persistent, somewhat thickened, dry petioles); not scapose. | Perennials; caudex simple or branched (branches short); not scapose. |
Stems | unbranched, 0.08–0.6 dm, densely to moderately pubescent throughout, trichomes simple and stalked, 2- or 3-rayed, 0.2–1 mm. |
unbranched or branched distally, (0.6–)1–3(–3.8) dm, pubescent throughout, trichomes simple, 0.5–1mm, with 4–10-rayed ones, 0.05–0.2 mm (mostly branched on basal parts). |
Basal leaves | rosulate; petiolate; petiole ciliate, (trichomes simple, 0.2–1 mm); blade linear to linear-oblanceolate, (0.4–)0.6–1.5(–2) cm × 1–2 mm, margins entire, (pubescent as petiole), surfaces pubescent abaxially with simple and stalked, 2- or 3-rayed trichomes, 0.1–0.4 mm, adaxially glabrate or subapically sparsely pubescent with simple trichomes. |
rosulate; petiolate; petiole ciliate throughout; blade linear to oblanceolate or oblong, (0.5–)0.8–2(–3.5) cm × 1.5–4(–11) mm, margins entire or dentate, (base and apex ciliate, trichomes simple, 0.3–0.8 mm), surfaces pubescent with short-stalked, 4–12-rayed trichomes, 0.1–0.3 mm. |
Cauline leaves | 1–4; sessile; blade linear to linear-oblanceolate, margins entire, surfaces pubescent as basal. |
3–10(–17), (not overlapping); sessile; blade lanceolate to ovate or oblong, margins entire or dentate, surfaces pubescent as basal (adaxially with simple and forked trichomes near blade base). |
Racemes | 5–12(–16)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
(10–)15–47(–63)-flowered, basally bracteate, often considerably elongated in fruit; rachis not flexuous, densely pubescent, trichomes 4–10-rayed (0.05–0.2 mm), and fewer simple ones. |
Flowers | sepals ovate, 1.5–2 mm, pubescent, (trichomes simple and short-stalked, 2- or 3-rayed); petals bright yellow, spatulate, 3–4.5 × 1–2 mm, (flared, clawed); anthers ovate, 0.3–0.5 mm. |
sepals (green or lavender), oblong to ovate, 1.5–2 mm, pubescent, (trichomes simple and few-rayed); petals white, oblanceolate to spatulate, 2.3–3.7(–4.5) × 0.7–1.7 mm; anthers ovate, 0.1–0.2 mm. |
Fruiting pedicels | divaricate-ascending, straight or slightly curved upward, 1.5–6 mm, pubescent as stem. |
suberect or ascending, straight, 2–5(–10) mm, pubescent as rachis. |
Fruits | lanceolate to oblong-lanceolate, slightly twisted or plane, flattened, 4–12 × 1.5–3 mm; valves glabrous or pubescent, trichomes simple, 0.07–0.2 mm; ovules 16–20 per ovary; style 0.4–1.2 mm. |
(subappressed to rachis), linear-lanceolate to linear or, rarely, ovate-oblong, slightly twisted or plane, flattened, (5–)6–11 × 1.5–2(–2.5) mm; valves pubescent, trichomes short-stalked, 3–7-rayed, 0.05–0.3 mm; ovules 28–48 per ovary; style 0.1–0.6 mm. |
Seeds | ovoid, 1–1.4 × 0.7–1 mm. |
ovoid, 0.5–0.7(–0.9) × 0.3–0.5 mm. |
2n | = 24 ± 3. |
= 32. |
Draba grayana |
Draba cana |
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Phenology | Flowering Jul–Aug. | Flowering (May-)Jun–Aug. |
Habitat | Alpine tundra, fellfields, gravelly wet meadows | Rock outcrops and talus, open prairie benchlands, roadsides, meadows, alpine tundra |
Elevation | 3500-4000 m (11500-13100 ft) | 0-4100 m (0-13500 ft) |
Distribution |
CO |
AK; CA; CO; ID; ME; MI; MT; NH; NM; NV; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NF; NT; ON; QC; SK; YT; Greenland
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Discussion | Of conservation concern. Draba grayana is an apomictic species clearly related to D. exunguiculata, with which it is occasionally sympatric (see discussion of 36. D. exunguiculata for distinguishing features). It is known from Chaffe, Clear Creek, Larimer, Park, and Summit counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The limits of Draba cana have long been confused, and the species was treated as a synonym of the Himalayan D. lanceolata Royle (M. L. Fernald 1934; C. L. Hitchcock 1941) or as a variety of the western North American D. breweri (R. C. Rollins 1993). However, G. A. Mulligan (1971) clearly demonstrated that all three are distinct and should be maintained. Some Utah plants corresponding to the type of D. valida have shorter and wider oblong-ovate fruits. In all other aspects, they are indistinguishable from D. cana. Additional studies are needed to establish whether such plants should be formally recognized. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 310. | FNA vol. 7, p. 298. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. streptocarpa var. grayana, D. alpicola | D. breweri var. cana, D. valida |
Name authority | (Rydberg) C. L. Hitchcock: Revis. Drabas W. N. Amer., 29. (1941) | Rydberg: Bull. Torrey Bot. Club 29: 241. (1902) |
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