Draba grandis |
Draba streptobrachia |
|
---|---|---|
north Pacific Draba |
alpine tundra Draba |
|
Habit | Perennials; caudex simple or branched (fleshy, with persistent leaf bases); not scapose. | Perennials; caudex branched (often with persistent leaf remains, branches not creeping); not scapose. |
Stems | (decumbent), unbranched, (0.2–)0.5–2.7(–3.7) dm, pubescent throughout, trichomes 2–4-rayed, 0.05–0.2 mm. |
unbranched, (0.1–)0.2–1(–1.3) dm, pubescent throughout, trichomes subsessile (often crisped), 3–5-rayed, stellate, 0.03–0.25(–0.4) mm, (rays sometimes forked). |
Basal leaves | rosulate; long-petiolate; petiole (winged, (1–)4–15 cm), often not ciliate (or ciliate to blade apex, trichomes simple, 0.3–0.8 mm); blade (somewhat fleshy), oblanceolate to spatulate or obovate, (1–)2–11(–17) cm × (5–)8–30(–45) mm, margins often coarsely dentate, (pubescent as petiole), surfaces usually pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.05–0.25 mm, adaxially similar, or also with simple and long-stalked, 2-rayed trichomes, to 0.8 mm, rarely glabrescent, with mostly simple and 2-rayed trichomes. |
rosulate; petiole (obscure), usually not ciliate, rarely sparsely pubescent, (trichomes simple, to 0.6 mm); blade oblanceolate to linear-oblanceolate, (0.4–)0.5–3(–4) cm × 1–5 mm, margins entire, surfaces pubescent with short-stalked (crisped), 3–8-rayed trichomes, 0.05–0.4 mm. |
Cauline leaves | 2–12(–16); sessile or petiolate; blade oblanceolate to obovate, margins dentate or entire, surfaces pubescent as basal. |
(1 or) 2–4 (or 5); sessile; blade oblong to ovate or linear, margins entire, surfaces pubescent as basal. |
Racemes | 5–26(–32)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
4–10(–18)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals broadly ovate, 3–4 mm, pubescent, (trichomes simple and short-stalked, 2-rayed); petals yellow, oblanceolate to spatulate, 4.5–7 × 1.8–3 mm; anthers oblong, 0.7–1 mm. |
sepals ovate, 2–3 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals yellow, spatulate, 3–5 × 1.5–3 mm, (clawed); anthers ovate, 0.25–0.4 mm. |
Fruiting pedicels | horizontal to divaricate-ascending or ascending, usually straight, rarely curved upward, (5–)10–22(–27) mm, pubescent, trichomes 2–4-rayed (0.05–0.3 mm), sometimes with simple and spurred ones. |
ascending, usually straight, rarely curved upward, (2–)3–8(–12) mm, pubescent as stem. |
Fruits | oblong to lanceolate, or ovate to suborbicular, slightly twisted or plane, flattened, (6–)10–20(–25) × 4–7(–9) mm; valves glabrous; ovules 24–52 per ovary; style (0.2–)0.4–1.6(–2) mm. |
(not appressed to rachis), ovate to elliptic or lanceolate, slightly twisted or plane, flattened, (3–)5–10 × 2–4 mm; valves often pubescent, occasionally glabrous, trichomes simple and minutely stalked, 2–4-rayed, 0.03–0.25 mm; ovules 10–16(–18) per ovary; style 0.3–0.8(–1.2) mm. |
Seeds | ovoid, 1.4–2 × 0.8–1.2 mm. |
oblong, 1–1.6 × 0.6–1 mm. |
2n | = 36. |
= ca. 64. |
Draba grandis |
Draba streptobrachia |
|
Phenology | Flowering May–Jul. | Flowering Jul–Aug. |
Habitat | Rocky bluffs above salt-water beaches, loamy seaside banks, sea bird rookeries, coastal herbaceous tundra and sandy blowouts | Alpine tundra, scree, ridges and alpine slopes, turf, fellfields, talus slopes, crevices in rock ledges, loose soils |
Elevation | 0-50(-200) m (0-200(-700) ft) | 3200-4000 m (10500-13100 ft) |
Distribution |
AK; BC; e Asia (Kuril and Ratmanov islands, Russian Far East) |
CO
|
Discussion | Almost all recent North American authors have used the name Draba hyperborea for this species. A. N. Berkutenko (1995) clearly showed that the type of that name belongs to an entirely different species that she placed in the genus Schivereckia Andrzejowski ex de Candolle. Draba grandis thus becomes the correct name for the North American taxon. Except for its fleshy leaves, Nesodraba is indistinguishable morphologically and molecularly from other species of Draba. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As indicated by Price, Draba streptobrachia is an apomict, yielding abundant, well-developed seed despite producing only abortive pollen. Morphological studies (M. D. Windham, unpubl.) suggest that the species may be an allopolyploid containing a genome from D. crassa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 309. | FNA vol. 7, p. 341. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Cochlearia siliquosa, Cochlearia spathulata, D. greenei, D. hatchiae, D. hyperborea var. spathulata, D. spathulata, Nesodraba grandis, Nesodraba megalocarpa, Nesodraba siliquosa | D. chrysantha, D. spectabilis var. dasycarpa |
Name authority | Langsdorff ex de Candolle: Syst. Nat. 2: 355. (1821) | R. A. Price: Brittonia 32: 168. (1980) |
Web links |
|