Draba grandis |
Draba pterosperma |
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north Pacific Draba |
wing-seed Draba, wingedseed Draba |
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Habit | Perennials; caudex simple or branched (fleshy, with persistent leaf bases); not scapose. | Perennials; (densely cespitose to pulvinate); caudex branched (with persistent leaves, branches sometimes terminating in sterile rosettes); scapose. |
Stems | (decumbent), unbranched, (0.2–)0.5–2.7(–3.7) dm, pubescent throughout, trichomes 2–4-rayed, 0.05–0.2 mm. |
unbranched, 0.3–1.1 dm, pubescent throughout, trichomes sub-dendritic, 2–7-rayed, (often crisped), 0.2–0.7 mm. |
Basal leaves | rosulate; long-petiolate; petiole (winged, (1–)4–15 cm), often not ciliate (or ciliate to blade apex, trichomes simple, 0.3–0.8 mm); blade (somewhat fleshy), oblanceolate to spatulate or obovate, (1–)2–11(–17) cm × (5–)8–30(–45) mm, margins often coarsely dentate, (pubescent as petiole), surfaces usually pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.05–0.25 mm, adaxially similar, or also with simple and long-stalked, 2-rayed trichomes, to 0.8 mm, rarely glabrescent, with mostly simple and 2-rayed trichomes. |
(densely imbricate); rosulate; petiolate; petiole base and margin ciliate, (trichomes simple or branched, often crisped, 0.3–1 mm); blade oblanceolate to obovate or spatulate, 0.2–0.7 cm × 1–2.8 mm, margins entire, surfaces densely pubescent (often grayish) with stellate-dendritic, stalked, 5–12-rayed, (often crisped) trichomes, 0.2–0.6 mm. |
Cauline leaves | 2–12(–16); sessile or petiolate; blade oblanceolate to obovate, margins dentate or entire, surfaces pubescent as basal. |
0. |
Racemes | 5–26(–32)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
4–12-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals broadly ovate, 3–4 mm, pubescent, (trichomes simple and short-stalked, 2-rayed); petals yellow, oblanceolate to spatulate, 4.5–7 × 1.8–3 mm; anthers oblong, 0.7–1 mm. |
sepals oblong, 3–4 mm, pubescent, (trichomes stalked, branched, crisped); petals yellow, oblanceolate, 6–7 × 1.5–2.5 mm; anthers oblong, 0.7–0.9 mm. |
Fruiting pedicels | horizontal to divaricate-ascending or ascending, usually straight, rarely curved upward, (5–)10–22(–27) mm, pubescent, trichomes 2–4-rayed (0.05–0.3 mm), sometimes with simple and spurred ones. |
erect (and subappressed to rachis) or ascending, straight, (3–)4–9(–12) mm, pubescent as stem. |
Fruits | oblong to lanceolate, or ovate to suborbicular, slightly twisted or plane, flattened, (6–)10–20(–25) × 4–7(–9) mm; valves glabrous; ovules 24–52 per ovary; style (0.2–)0.4–1.6(–2) mm. |
broadly ovate to lanceolate or subelliptic, plane, flattened, 5–10(–13) × (2.5–)3.5–5(–6) mm; valves pubescent, trichomes short stalked, 2–7-rayed, (straight or crisped), 0.15–0.6 mm; ovules 8–12 per ovary; style 1.5–3(–3.8) mm. |
Seeds | ovoid, 1.4–2 × 0.8–1.2 mm. |
(winged), ovate, 1.6–3 × 1.3–1.8 mm; (wing 0.3–0.9 mm wide). |
2n | = 36. |
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Draba grandis |
Draba pterosperma |
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Phenology | Flowering May–Jul. | Flowering Jun–Sep. |
Habitat | Rocky bluffs above salt-water beaches, loamy seaside banks, sea bird rookeries, coastal herbaceous tundra and sandy blowouts | Limestone and marble outcrops, talus, gravel slopes |
Elevation | 0-50(-200) m (0-200(-700) ft) | 1500-2500 m (4900-8200 ft) |
Distribution |
AK; BC; e Asia (Kuril and Ratmanov islands, Russian Far East) |
CA |
Discussion | Almost all recent North American authors have used the name Draba hyperborea for this species. A. N. Berkutenko (1995) clearly showed that the type of that name belongs to an entirely different species that she placed in the genus Schivereckia Andrzejowski ex de Candolle. Draba grandis thus becomes the correct name for the North American taxon. Except for its fleshy leaves, Nesodraba is indistinguishable morphologically and molecularly from other species of Draba. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba pterosperma is a distinctive species that is restricted to the Marble Mountains of northern California (Siskiyou County). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 309. | FNA vol. 7, p. 332. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Cochlearia siliquosa, Cochlearia spathulata, D. greenei, D. hatchiae, D. hyperborea var. spathulata, D. spathulata, Nesodraba grandis, Nesodraba megalocarpa, Nesodraba siliquosa | |
Name authority | Langsdorff ex de Candolle: Syst. Nat. 2: 355. (1821) | Payson: Amer. J. Bot. 4: 266. (1917) |
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