FNA: Draba grandis vs. Draba lemmonii

	Draba grandis	Draba lemmonii
	north Pacific Draba	granite Draba, Lemmon's Draba
Habit	Perennials; caudex simple or branched (fleshy, with persistent leaf bases); not scapose.	Perennials; (cespitose, not pulvinate); caudex branched (covered with persistent petiole remains, branches sometimes terminating in sterile rosettes); scapose.
Stems	(decumbent), unbranched, (0.2–)0.5–2.7(–3.7) dm, pubescent throughout, trichomes 2–4-rayed, 0.05–0.2 mm.	unbranched, 0.3–1(–1.5) dm, hirsute throughout, trichomes simple, 0.2–0.7 mm, with short-stalked, 2-rayed ones, (smaller).
Basal leaves	rosulate; long-petiolate; petiole (winged, (1–)4–15 cm), often not ciliate (or ciliate to blade apex, trichomes simple, 0.3–0.8 mm); blade (somewhat fleshy), oblanceolate to spatulate or obovate, (1–)2–11(–17) cm × (5–)8–30(–45) mm, margins often coarsely dentate, (pubescent as petiole), surfaces usually pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.05–0.25 mm, adaxially similar, or also with simple and long-stalked, 2-rayed trichomes, to 0.8 mm, rarely glabrescent, with mostly simple and 2-rayed trichomes.	rosulate; shortly petiolate; petiole base and margin ciliate, (trichomes simple, 0.5–1 mm, midvein obscure); blade (somewhat fleshy), oblanceolate to obovate, 0.4–1(–1.8) cm × 1.5–4(–6) mm, margins entire, surfaces hirsute, abaxially mostly with stalked, 2-rayed trichomes, 0.1–0.8 mm, rarely with fewer, simple ones, adaxially with mostly simple ones, 0.5–1 mm.
Cauline leaves	2–12(–16); sessile or petiolate; blade oblanceolate to obovate, margins dentate or entire, surfaces pubescent as basal.	0.
Racemes	5–26(–32)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem.	4–15(–21)-flowered, ebracteate, elongated in fruit; rachis not flexuous, hirsute as stem.
Flowers	sepals broadly ovate, 3–4 mm, pubescent, (trichomes simple and short-stalked, 2-rayed); petals yellow, oblanceolate to spatulate, 4.5–7 × 1.8–3 mm; anthers oblong, 0.7–1 mm.	sepals ovate, 2–2.7 mm, pubescent, (trichomes simple, short-stalked, 2-rayed); petals yellow, spatulate, 4–6 × 1.5–2.5 mm; anthers oblong, 0.4–0.6 mm.
Fruiting pedicels	horizontal to divaricate-ascending or ascending, usually straight, rarely curved upward, (5–)10–22(–27) mm, pubescent, trichomes 2–4-rayed (0.05–0.3 mm), sometimes with simple and spurred ones.	horizontal to divaricate-ascending (often somewhat decurrent basally), straight or curved upward, 4–10(–14) mm, hirsute as stem.
Fruits	oblong to lanceolate, or ovate to suborbicular, slightly twisted or plane, flattened, (6–)10–20(–25) × 4–7(–9) mm; valves glabrous; ovules 24–52 per ovary; style (0.2–)0.4–1.6(–2) mm.	ovate to ovate-lanceolate, slightly twisted, flattened, 4–9 × 3.5–5 mm; valves pubescent, trichomes usually simple, 0.1–0.45 mm, rarely with fewer, short-stalked, 2-rayed ones; ovules 10–14(–16) per ovary; style 0.1–0.6(–0.8) mm.
Seeds	ovoid, 1.4–2 × 0.8–1.2 mm.	ovoid, 1–1.4 × 0.6–1 mm.
2n	= 36.	= 50.

	Draba grandis	Draba lemmonii
Phenology	Flowering May–Jul.	Flowering Jul–Aug.
Habitat	Rocky bluffs above salt-water beaches, loamy seaside banks, sea bird rookeries, coastal herbaceous tundra and sandy blowouts	Granitic rock outcrops, boulder slopes, alpine fellfields
Elevation	0-50(-200) m (0-200(-700) ft)	3000-4000 m (9800-13100 ft)
Distribution	AK; BC; e Asia (Kuril and Ratmanov islands, Russian Far East) [BONAP county map]	CA [WildflowerSearch map] [BONAP county map]
Discussion	Almost all recent North American authors have used the name Draba hyperborea for this species. A. N. Berkutenko (1995) clearly showed that the type of that name belongs to an entirely different species that she placed in the genus Schivereckia Andrzejowski ex de Candolle. Draba grandis thus becomes the correct name for the North American taxon. Except for its fleshy leaves, Nesodraba is indistinguishable morphologically and molecularly from other species of Draba. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Draba lemmonii was so broadly circumscribed by C. L. Hitchcock (1941) and R. C. Rollins (1993) that it included plants here assigned to three different species. For a list of features distinguishing D. lemmonii from the recently-segregated D. longisquamosa and D. cyclomorpha, see I. A. Al-Shehbaz and M. D. Windham (2007). Draba lemmonii is apparently restricted to alpine areas of the Sierra Nevada in Fresno, Inyo, Madera, Mono, and Tuolumne counties. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 7, p. 309.	FNA vol. 7, p. 316.
Parent taxa	Brassicaceae > tribe Arabideae > Draba	Brassicaceae > tribe Arabideae > Draba
Sibling taxa	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms	Cochlearia siliquosa, Cochlearia spathulata, D. greenei, D. hatchiae, D. hyperborea var. spathulata, D. spathulata, Nesodraba grandis, Nesodraba megalocarpa, Nesodraba siliquosa
Name authority	Langsdorff ex de Candolle: Syst. Nat. 2: 355. (1821)	S. Watson: in W. H. Brewer et al., Bot. California 2: 430. (1880)
Web links	Local floras: BC Local Web sites: PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA, OR Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Draba lemmonii

north Pacific Draba

granite Draba, Lemmon's Draba

Habit

Perennials; caudex simple or branched (fleshy, with persistent leaf bases); not scapose.

Perennials; (cespitose, not pulvinate); caudex branched (covered with persistent petiole remains, branches sometimes terminating in sterile rosettes); scapose.

Stems

(decumbent), unbranched, (0.2–)0.5–2.7(–3.7) dm, pubescent throughout, trichomes 2–4-rayed, 0.05–0.2 mm.

unbranched, 0.3–1(–1.5) dm, hirsute throughout, trichomes simple, 0.2–0.7 mm, with short-stalked, 2-rayed ones, (smaller).

Basal leaves

rosulate; long-petiolate;

petiole (winged, (1–)4–15 cm), often not ciliate (or ciliate to blade apex, trichomes simple, 0.3–0.8 mm);

blade (somewhat fleshy), oblanceolate to spatulate or obovate, (1–)2–11(–17) cm × (5–)8–30(–45) mm, margins often coarsely dentate, (pubescent as petiole), surfaces usually pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.05–0.25 mm, adaxially similar, or also with simple and long-stalked, 2-rayed trichomes, to 0.8 mm, rarely glabrescent, with mostly simple and 2-rayed trichomes.

rosulate; shortly petiolate;

petiole base and margin ciliate, (trichomes simple, 0.5–1 mm, midvein obscure);

blade (somewhat fleshy), oblanceolate to obovate, 0.4–1(–1.8) cm × 1.5–4(–6) mm, margins entire, surfaces hirsute, abaxially mostly with stalked, 2-rayed trichomes, 0.1–0.8 mm, rarely with fewer, simple ones, adaxially with mostly simple ones, 0.5–1 mm.

Cauline leaves

2–12(–16);

sessile or petiolate;

blade oblanceolate to obovate, margins dentate or entire, surfaces pubescent as basal.

Racemes

5–26(–32)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

4–15(–21)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, hirsute as stem.

Flowers

sepals broadly ovate, 3–4 mm, pubescent, (trichomes simple and short-stalked, 2-rayed);

petals yellow, oblanceolate to spatulate, 4.5–7 × 1.8–3 mm;

anthers oblong, 0.7–1 mm.

sepals ovate, 2–2.7 mm, pubescent, (trichomes simple, short-stalked, 2-rayed);

petals yellow, spatulate, 4–6 × 1.5–2.5 mm;

anthers oblong, 0.4–0.6 mm.

Fruiting pedicels

horizontal to divaricate-ascending or ascending, usually straight, rarely curved upward, (5–)10–22(–27) mm, pubescent, trichomes 2–4-rayed (0.05–0.3 mm), sometimes with simple and spurred ones.

horizontal to divaricate-ascending (often somewhat decurrent basally), straight or curved upward, 4–10(–14) mm, hirsute as stem.

Fruits

oblong to lanceolate, or ovate to suborbicular, slightly twisted or plane, flattened, (6–)10–20(–25) × 4–7(–9) mm;

valves glabrous;

ovules 24–52 per ovary;

style (0.2–)0.4–1.6(–2) mm.

ovate to ovate-lanceolate, slightly twisted, flattened, 4–9 × 3.5–5 mm;

valves pubescent, trichomes usually simple, 0.1–0.45 mm, rarely with fewer, short-stalked, 2-rayed ones;

ovules 10–14(–16) per ovary;

style 0.1–0.6(–0.8) mm.

Seeds

ovoid, 1.4–2 × 0.8–1.2 mm.

ovoid, 1–1.4 × 0.6–1 mm.

= 36.

= 50.

Draba grandis

Draba lemmonii

Phenology

Flowering May–Jul.

Flowering Jul–Aug.

Habitat

Rocky bluffs above salt-water beaches, loamy seaside banks, sea bird rookeries, coastal herbaceous tundra and sandy blowouts

Granitic rock outcrops, boulder slopes, alpine fellfields

Elevation

0-50(-200) m (0-200(-700) ft)

3000-4000 m (9800-13100 ft)

Distribution

AK; BC; e Asia (Kuril and Ratmanov islands, Russian Far East)

[BONAP county map]

[WildflowerSearch map]

[BONAP county map]

Discussion

Almost all recent North American authors have used the name Draba hyperborea for this species. A. N. Berkutenko (1995) clearly showed that the type of that name belongs to an entirely different species that she placed in the genus Schivereckia Andrzejowski ex de Candolle. Draba grandis thus becomes the correct name for the North American taxon. Except for its fleshy leaves, Nesodraba is indistinguishable morphologically and molecularly from other species of Draba.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba lemmonii was so broadly circumscribed by C. L. Hitchcock (1941) and R. C. Rollins (1993) that it included plants here assigned to three different species. For a list of features distinguishing D. lemmonii from the recently-segregated D. longisquamosa and D. cyclomorpha, see I. A. Al-Shehbaz and M. D. Windham (2007). Draba lemmonii is apparently restricted to alpine areas of the Sierra Nevada in Fresno, Inyo, Madera, Mono, and Tuolumne counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 7, p. 309.

FNA vol. 7, p. 316.

Parent taxa

Brassicaceae > tribe Arabideae > Draba

Sibling taxa

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

Synonyms

Cochlearia siliquosa, Cochlearia spathulata, D. greenei, D. hatchiae, D. hyperborea var. spathulata, D. spathulata, Nesodraba grandis, Nesodraba megalocarpa, Nesodraba siliquosa

Name authority

Langsdorff ex de Candolle: Syst. Nat. 2: 355. (1821)

S. Watson: in W. H. Brewer et al., Bot. California 2: 430. (1880)

Web links

Local floras: CA, OR
Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora

The species comparison tool is brought to you by:

Flora of North America species comparison

Draba grandis

Draba lemmonii

Draba grandis

Draba lemmonii