FNA: Draba graminea vs. Draba santaquinensis

	Draba graminea	Draba santaquinensis
	Rocky Mountain Draba
Habit	Perennials; (cespitose); caudex simple or branched (with persistent leaf bases); not scapose.	Biennials or perennials; (short-lived); caudex simple (poorly developed, without persistent leaf bases); not scapose.
Stems	unbranched, 0.1–0.5(–0.8) dm, usually pubescent throughout, rarely glabrous, trichomes simple or subsessile, 2-rayed or spurred, (crisped), 0.1–0.5 mm.	usually branched, 1.1–3.4 dm, hirsute proximally, trichomes mostly simple and 2-rayed, 0.2–1.5 mm, pubescent distally, trichomes mostly 2- or 3-rayed, 0.2–1 mm.
Basal leaves	rosulate; petiolate; petiole ciliate, (trichomes usually straight and simple, rarely also 2-rayed, 0.2–0.6(–0.8) mm); blade linear to linear-oblanceolate, (0.5–)1–4 cm × 0.3–2(–3) mm, margins entire (pubescent as petiole), surfaces glabrous.	rosulate; petiole (obscure), ciliate, (trichomes simple and 2-rayed); blade obovate to oblanceolate, 1.6–2.5 cm × 5–10 mm, margins usually entire, rarely denticulate, surfaces pubescent, abaxially with stalked, mostly cruciform trichomes, 0.5–1 mm, adaxially with simple and stalked, 2–4-rayed ones.
Cauline leaves	(1–)3–9(–12) (as bracts); sessile; blade linear to oblanceolate or lanceolate, margins entire, (ciliate proximally, similar to basal).	usually 1–5; sessile or subsessile; blade oblanceolate to oblong, margins often denticulate, (ciliate proximally, with 2- or 3-rayed trichomes).
Racemes	3–15-flowered, bracteate, elongated in fruit; rachis not flexuous, usually pubescent as stem, rarely glabrous.	9–25-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent, trichomes 2- or 3-rayed.
Flowers	sepals ovate, 1.5–2.5 mm, glabrous or pubescent, (trichomes simple and short-stalked, 2-rayed); petals yellow, spatulate to obovate, 3–5 × 1.5–3 mm; anthers ovate, 0.3–0.5 mm.	sepals oblong-obovate, 2–3.5 mm, pubescent, (trichomes short- to long-stalked, 2–4-rayed, 0.2–0.8 mm); petals yellow (often fading whitish), oblanceolate, 3–5 × 1–1.5 mm; anthers ovate, 0.4–0.5 mm.
Fruiting pedicels	divaricate-ascending, usually straight, rarely curved upward, 3–10(–15) mm, usually pubescent as stem, rarely glabrous.	horizontal to divaricate-ascending, usually straight, rarely curved upward, 7–10 mm, pubescent, trichomes 2- or 3-rayed.
Fruits	ovate-elliptic to lanceolate, slightly twisted or plane, flattened, 5–11 × 2.5–5 mm; valves glabrous; ovules 8–16 per ovary; style 0.2–0.7 mm.	narrowly elliptic to slightly falcate, plane, flattened, (7–)10–16 × (2–)2.5–3.7 mm; valves pubescent, trichomes short-stalked, 2–4-rayed, 0.2–0.5 mm; ovules 14–22 per ovary; style 0.9–1.2 mm.
Seeds	ovoid, 1.2–1.5 × 0.7–1 mm.	oblong, 1.1–1.5 × 0.7–0.9 mm.
2n	= 18.	= 20.

	Draba graminea	Draba santaquinensis
Phenology	Flowering Jul–Sep.	Flowering late Apr-early Jun.
Habitat	Rocky areas and ridges, alpine tundra, gravel bars in streams	Limestone outcrops and rocky slopes in mixed conifer communities
Elevation	3000-4100 m (9800-13500 ft)	1800-2400 m (5900-7900 ft)
Distribution	CO [WildflowerSearch map] [BONAP county map]	UT [BONAP county map]
Discussion	Of conservation concern. Draba graminea is occasionally confused with D. crassa, which occupies similar habitats and elevations. It is easily distinguished from the latter by having narrower [0.3–2(–3) versus 2.5–8(–10) mm wide] basal leaves and bracteate (versus ebracteate) racemes. It is known to us only from Hinsdale, La Plata, Ouray, San Juan, and San Miguel counties. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Although Draba santaquinensis was included within D. brachystylis by previous authors, I. A. Al-Shehbaz and M. D. Windham (2007) have shown that it is distinct both morphologically and chromosomally. It is currently known only from Utah County (American Fork, Provo, and Santaquin canyons) in north-central Utah. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 7, p. 309.	FNA vol. 7, p. 334.
Parent taxa	Brassicaceae > tribe Arabideae > Draba	Brassicaceae > tribe Arabideae > Draba
Sibling taxa	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms	Braya graminea, D. chrysantha var. graminea, D. chrysantha var. hirticaulis
Name authority	Greene: Pl. Baker. 3: 5. (1901)	Windham & Allphin: Harvard Pap. Bot. 12: 410. (2007)
Web links	Local Web sites: SW CO Wildflowers WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Draba santaquinensis

Rocky Mountain Draba

Habit

Perennials; (cespitose); caudex simple or branched (with persistent leaf bases); not scapose.

Biennials or perennials; (short-lived); caudex simple (poorly developed, without persistent leaf bases); not scapose.

Stems

unbranched, 0.1–0.5(–0.8) dm, usually pubescent throughout, rarely glabrous, trichomes simple or subsessile, 2-rayed or spurred, (crisped), 0.1–0.5 mm.

usually branched, 1.1–3.4 dm, hirsute proximally, trichomes mostly simple and 2-rayed, 0.2–1.5 mm, pubescent distally, trichomes mostly 2- or 3-rayed, 0.2–1 mm.

Basal leaves

rosulate; petiolate;

petiole ciliate, (trichomes usually straight and simple, rarely also 2-rayed, 0.2–0.6(–0.8) mm);

blade linear to linear-oblanceolate, (0.5–)1–4 cm × 0.3–2(–3) mm, margins entire (pubescent as petiole), surfaces glabrous.

rosulate;

petiole (obscure), ciliate, (trichomes simple and 2-rayed);

blade obovate to oblanceolate, 1.6–2.5 cm × 5–10 mm, margins usually entire, rarely denticulate, surfaces pubescent, abaxially with stalked, mostly cruciform trichomes, 0.5–1 mm, adaxially with simple and stalked, 2–4-rayed ones.

Cauline leaves

(1–)3–9(–12) (as bracts);

sessile;

blade linear to oblanceolate or lanceolate, margins entire, (ciliate proximally, similar to basal).

usually 1–5;

sessile or subsessile;

blade oblanceolate to oblong, margins often denticulate, (ciliate proximally, with 2- or 3-rayed trichomes).

Racemes

3–15-flowered, bracteate, elongated in fruit;

rachis not flexuous, usually pubescent as stem, rarely glabrous.

9–25-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent, trichomes 2- or 3-rayed.

Flowers

sepals ovate, 1.5–2.5 mm, glabrous or pubescent, (trichomes simple and short-stalked, 2-rayed);

petals yellow, spatulate to obovate, 3–5 × 1.5–3 mm;

anthers ovate, 0.3–0.5 mm.

sepals oblong-obovate, 2–3.5 mm, pubescent, (trichomes short- to long-stalked, 2–4-rayed, 0.2–0.8 mm);

petals yellow (often fading whitish), oblanceolate, 3–5 × 1–1.5 mm;

anthers ovate, 0.4–0.5 mm.

Fruiting pedicels

divaricate-ascending, usually straight, rarely curved upward, 3–10(–15) mm, usually pubescent as stem, rarely glabrous.

horizontal to divaricate-ascending, usually straight, rarely curved upward, 7–10 mm, pubescent, trichomes 2- or 3-rayed.

Fruits

ovate-elliptic to lanceolate, slightly twisted or plane, flattened, 5–11 × 2.5–5 mm;

valves glabrous;

ovules 8–16 per ovary;

style 0.2–0.7 mm.

narrowly elliptic to slightly falcate, plane, flattened, (7–)10–16 × (2–)2.5–3.7 mm;

valves pubescent, trichomes short-stalked, 2–4-rayed, 0.2–0.5 mm;

ovules 14–22 per ovary;

style 0.9–1.2 mm.

Seeds

ovoid, 1.2–1.5 × 0.7–1 mm.

oblong, 1.1–1.5 × 0.7–0.9 mm.

= 18.

= 20.

Draba graminea

Draba santaquinensis

Phenology

Flowering Jul–Sep.

Flowering late Apr-early Jun.

Habitat

Rocky areas and ridges, alpine tundra, gravel bars in streams

Limestone outcrops and rocky slopes in mixed conifer communities

Elevation

3000-4100 m (9800-13500 ft)

1800-2400 m (5900-7900 ft)

Distribution

[WildflowerSearch map]

[BONAP county map]

Discussion

Of conservation concern.

Draba graminea is occasionally confused with D. crassa, which occupies similar habitats and elevations. It is easily distinguished from the latter by having narrower [0.3–2(–3) versus 2.5–8(–10) mm wide] basal leaves and bracteate (versus ebracteate) racemes. It is known to us only from Hinsdale, La Plata, Ouray, San Juan, and San Miguel counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although Draba santaquinensis was included within D. brachystylis by previous authors, I. A. Al-Shehbaz and M. D. Windham (2007) have shown that it is distinct both morphologically and chromosomally. It is currently known only from Utah County (American Fork, Provo, and Santaquin canyons) in north-central Utah.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 7, p. 309.

FNA vol. 7, p. 334.

Parent taxa

Brassicaceae > tribe Arabideae > Draba

Sibling taxa

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

Synonyms

Braya graminea, D. chrysantha var. graminea, D. chrysantha var. hirticaulis

Name authority

Greene: Pl. Baker. 3: 5. (1901)

Windham & Allphin: Harvard Pap. Bot. 12: 410. (2007)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Draba graminea

Draba santaquinensis

Draba graminea

Draba santaquinensis