Draba graminea |
Draba lonchocarpa |
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Rocky Mountain Draba |
lance-fruit Draba, lance-fruit whitlow-grass, lance-pod Draba, lance-pod whitlow-grass, lancefruit Draba whitlow-wort, spear-fruit Draba |
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Habit | Perennials; (cespitose); caudex simple or branched (with persistent leaf bases); not scapose. | Perennials; (cespitose, sometimes forming mats); caudex branched (covered with persistent leaf bases); often scapose. |
Stems | unbranched, 0.1–0.5(–0.8) dm, usually pubescent throughout, rarely glabrous, trichomes simple or subsessile, 2-rayed or spurred, (crisped), 0.1–0.5 mm. |
unbranched, (0.1–)0.3–1.1 dm, glabrous throughout or pubescent proximally or throughout, trichomes minutely stalked, 8–12-rayed, (non-crisped), 0.08–0.3 mm. |
Basal leaves | rosulate; petiolate; petiole ciliate, (trichomes usually straight and simple, rarely also 2-rayed, 0.2–0.6(–0.8) mm); blade linear to linear-oblanceolate, (0.5–)1–4 cm × 0.3–2(–3) mm, margins entire (pubescent as petiole), surfaces glabrous. |
rosulate; shortly petiolate; petiole (obsolete), margin ciliate proximally, (trichomes soft, simple, 2-rayed, 0.2–0.6 mm); blade oblanceolate to obovate, (0.2–)0.3–1.5 cm × 1–3(–5) mm, margins entire, surfaces pubescent abaxially with short-stalked, stellate, 8–12-rayed, (non-crisped) trichomes, 0.15–0.4 mm, (midvein obscure), adaxially glabrous basally or with simple and long-stalked, branched trichomes. |
Cauline leaves | (1–)3–9(–12) (as bracts); sessile; blade linear to oblanceolate or lanceolate, margins entire, (ciliate proximally, similar to basal). |
0 or 1(–4); sessile; blade ovate or oblong, margins entire, surfaces pubescent as basal. |
Racemes | 3–15-flowered, bracteate, elongated in fruit; rachis not flexuous, usually pubescent as stem, rarely glabrous. |
3–9-flowered, usually ebracteate, rarely proximalmost 1(–4) flowers bracteate, elongated in fruit; rachis often slightly flexuous (in fruit), glabrous or pubescent as stem. |
Flowers | sepals ovate, 1.5–2.5 mm, glabrous or pubescent, (trichomes simple and short-stalked, 2-rayed); petals yellow, spatulate to obovate, 3–5 × 1.5–3 mm; anthers ovate, 0.3–0.5 mm. |
sepals ovate, 1.5–2 mm, pubescent, (trichomes simple and short-stalked, 2–5-rayed); petals white, oblanceolate, 2–3.5 × 1–1.5 mm; anthers ovate, 0.2–0.3 mm. |
Fruiting pedicels | divaricate-ascending, usually straight, rarely curved upward, 3–10(–15) mm, usually pubescent as stem, rarely glabrous. |
ascending, usually straight, rarely curved upward, 2–9(–15) mm, glabrous or pubescent as stem. |
Fruits | ovate-elliptic to lanceolate, slightly twisted or plane, flattened, 5–11 × 2.5–5 mm; valves glabrous; ovules 8–16 per ovary; style 0.2–0.7 mm. |
linear to narrowly lanceolate or narrowly oblong, slightly twisted or plane, flattened, 6–15(–18) × 1–2(–3) mm; valves glabrous or sparsely puberulent, trichomes simple and minutely stalked, 2-rayed, 0.1–0.2 mm; ovules 16–24(–28) per ovary; style (0.05–)0.1–0.25(–0.35) mm. |
Seeds | ovoid, 1.2–1.5 × 0.7–1 mm. |
ovoid, 0.7–1 × 0.5–0.6 mm. |
2n | = 18. |
= 16. |
Draba graminea |
Draba lonchocarpa |
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Phenology | Flowering Jul–Sep. | Flowering Jun–Jul. |
Habitat | Rocky areas and ridges, alpine tundra, gravel bars in streams | Rocky outcrops and ridges, loose talus, tundra |
Elevation | 3000-4100 m (9800-13500 ft) | (300-)2800-4000 m ((1000-)9200-13100 ft) |
Distribution |
CO
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AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; NT; YT; e Asia (Russian Far East, Siberia)
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Discussion | Of conservation concern. Draba graminea is occasionally confused with D. crassa, which occupies similar habitats and elevations. It is easily distinguished from the latter by having narrower [0.3–2(–3) versus 2.5–8(–10) mm wide] basal leaves and bracteate (versus ebracteate) racemes. It is known to us only from Hinsdale, La Plata, Ouray, San Juan, and San Miguel counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba lonchocarpa is a highly variable species within which O. E. Schulz (1927), G. A. Mulligan (1974), and R. C. Rollins (1993) recognized three to five varieties. By contrast, C. L. Hitchcock (1941) united it with D. nivalis and recognized six varieties (see 68. D. nivalis for differences). Some of the infraspecific taxa of D. lonchocarpa are based on trivial characteristics and are listed in the synonymy above without further comment. The most problematic are briefly discussed below. Authors recognizing var. vestita claim that it differs from var. lonchocarpa by having pubescent (versus glabrous) stems and pedicels, 1- or 2-leaved (versus 0 or 1-leaved) scapes, and fruits appressed (versus not appressed) to the rachises. These characteristics do not appear to be strongly correlated. A case in point is the holotype sheet of var. semitonsa, which includes plants with puberulent or glabrous fruits, as well as with pubescent and glabrous stems that are 0–4-leaved. Leafless and densely pubescent scapes are found in Trelease 3913 (MO), whereas completely glabrous, 0–2-leaved stems, and fully appressed fruits are found in Calder 5617a (DAO). Other exceptions can be cited, though the vast majority of the plants examined have leafless, glabrous scapes. An examination of the type collections of var. thompsonii, Thompson 9512 (holotype, UC; isotypes, DS, GH, MO, NY, RSA, US), clearly shows that the taxon usually has oblong to lanceolate fruits 2–3.2 mm wide, as opposed to linear fruits less than 2 mm wide in var. lonchocarpa. Indeed, a casual observation would immediately justify the recognition of var. thompsonii. Both fruit types can be found in plants of the same population (e.g., the RSA isotype) or even on the same plant (e.g., Thompson 10816, MO). Furthermore, fruits to 2.5 mm wide occur sporadically in various parts of the species range. For these reasons, and in the absence of a comprehensive study of the species, we choose to not recognize var. thompsonii at present. Draba lonchocarpa is found at elevations of 300–1200 m in Alaska. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 309. | FNA vol. 7, p. 317. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | Braya graminea, D. chrysantha var. graminea, D. chrysantha var. hirticaulis | D. nivalis var. elongata, D. lonchocarpa var. denudata, D. lonchocarpa var. exigua, D. lonchocarpa var. semitonsa, D. lonchocarpa var. thompsonii, D. lonchocarpa var. vestita, D. nivalis var. denudata, D. nivalis var. exigua, D. nivalis subsp. lonchocarpa, D. nivalis var. thompsonii |
Name authority | Greene: Pl. Baker. 3: 5. (1901) | Rydberg: Mem. New York Bot. Gard. 1: 181. (1900) |
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