Draba fladnizensis |
Draba longisquamosa |
|
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arctic Draba, Austrian Draba, Austrian whitlow-grass |
granite Draba |
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Habit | Perennials; (sometimes cespitose); caudex simple or branched (with persistent leaf bases); usually scapose. | Perennials; (cespitose); caudex branched (covered with persistent petiole remains, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, (0.2–)0.3–1(–1.3) dm, glabrous. |
unbranched, 0.2–0.9 dm, hirsute throughout, trichomes simple, 0.2–0.8 mm. |
Basal leaves | rosulate; petiole (obscure), margin ciliate, (trichomes simple or 2-rayed, 0.25–0.6 mm); blade linear to oblanceolate or narrowly obovate, (0.3–)0.4–1.2(–1.6) cm × 1–3(–4) mm, margins usually entire, rarely toothed, surfaces abaxially pubescent or glabrous, trichomes simple, sometimes with fewer, short-stalked, 2-rayed ones, (midvein prominent), adaxially often glabrous. |
rosulate; shortly petiolate; petiole base and margin ciliate, (trichomes simple, 0.1–1 mm); blade oblanceolate to obovate, 0.5–2 cm × 2–5 mm, margins entire, surfaces pubescent with simple trichomes, 0.15–0.8 mm. |
Cauline leaves | 0–2; sessile; blade oblong to ovate, margins entire, (ciliate). |
0. |
Racemes | (2 or) 3–11(–14)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous. |
4–16-flowered, ebracteate, (subumbellate), slightly elongated in fruit; rachis not flexuous, hirsute as stem. |
Flowers | sepals (green or purplish), ovate, 1.2–2.2 mm, glabrous or pubescent, (trichomes simple); petals white, spatulate, 2–2.5 × 0.8–1.5 mm; anthers ovate, 0.2–0.25 mm. |
sepals (persistent) ovate, 1.5–2.2 mm, pubescent, (trichomes simple); petals (persistent), yellow, spatulate, 3.5–5 × 1.5–2 mm; anthers oblong, 0.5–0.6 mm. |
Fruiting pedicels | divaricate-ascending, often straight, (1–)2–5(–6) mm, glabrous. |
divaricate to ascending (not decurrent basally), straight, 3–7 mm, hirsute as stem. |
Fruits | elliptic-lanceolate to oblong, plane, flattened, 3–8(–9) × 1.5–2 mm; valves glabrous; ovules 12–24 per ovary; style 0.05–0.2(–0.3) mm. |
ovate to suborbicular, plane, flattened, 3.5–7 × 2.5–5 mm; valves pubescent, trichomes simple, 0.1–0.3 mm; ovules 10–16 per ovary; style 0.4–1 mm. |
Seeds | oblong to elliptic, 0.8–1 × 0.5–0.6 mm. |
ovoid, 1–1.2 × 0.7–0.8 mm. |
2n | = 16. |
|
Draba fladnizensis |
Draba longisquamosa |
|
Phenology | Flowering Jun–Aug. | Flowering Jul. |
Habitat | Rock outcrops and talus, alpine meadows, sandy gravel | Gravelly areas |
Elevation | 0-1400 m at higher latitudes, 3000-3800 m at lower latitudes (0-4600 ft at higher latitudes, 9800-12500 ft at lower latitudes) | 3000-3900 m (9800-12800 ft) |
Distribution |
AK; CO; UT; WY; BC; NT; NU; QC; YT; Greenland; c Europe; s Europe; Asia; circumpolar and high alpine areas
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CA |
Discussion | Rollins reduced Draba pattersonii to a variety of D. fladnizensis and separated the two primarily on plant size and minor differences in fruit shape. Examination of D. fladnizensis specimens collected throughout Europe and North America reveals that the alleged differences between the two taxa are artificial. The type material of D. pattersonii, which was collected in Colorado, is a mixture of plants highly variable in their type of indumentum. The specimens have no flowers, but the habit, fruits, and leaves are nearly indistinguishable from those of D. fladnizensis from higher latitudes. In the absence of flowers, the white-flowered Draba fladnizensis (2n = 16) is often confused with the yellow-flowered D. crassifolia (2n = 40). The latter is an annual or short-lived perennial that rarely forms a well-developed caudex, whereas D. fladnizensis almost always produces a distinct caudex. Although most individuals of both species are scapose, they occasionally produce one or two cauline leaves. The cauline leaves are usually glabrous in D. crassifolia and ciliate in D. fladnizensis; in the latter, the distalmost cauline leaf usually subtends the proximalmost flower. Finally, the seeds in D. fladnizensis are slightly larger (0.8–0.1 × 0.5–0.6 versus 0.7–0.8 × 0.4–0.5 mm) than those of D. crassifolia, though the reliability of this distinction needs to be examined in greater detail. N. H. Holmgren (2005b) reported D. fladnizensis from central Nevada, but we have not seen any material of the species from that state. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba longisquamosa was treated as a synonym of D. lemmonii by both C. L. Hitchcock (1941) and R. C. Rollins (1993). We feel that the two taxa show sufficient morphological divergence and merit recognition as separate species (see I. A. Al-Shehbaz and M. D. Windham 2007 for more detailed discussion). Draba longisquamosa is currently known only from the southern Sierra Nevada in Fresno, Inyo, and Tulare counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 307. | FNA vol. 7, p. 317. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. fladnizensis var. pattersonii, D. pattersonii, D. pattersonii var. hirticaulis, D. wahlenbergii | |
Name authority | Wulfen: in N. J. Jacquin, Misc. Austriac. 1: 147, plate 17, fig. 1. (1778) | O. E. Schulz: in H. G. A. Engler, Pflanzenr. 89[IV,105]: 94. (1927) |
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