Draba fladnizensis |
Draba cana |
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arctic Draba, Austrian Draba, Austrian whitlow-grass |
Brewer's Draba, canescent Draba, canescent whitlow-mustard, cushion Draba, hoary Draba, hoary whitlow-grass, lance-leaf Draba, lance-leaf whitlow-grass |
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Habit | Perennials; (sometimes cespitose); caudex simple or branched (with persistent leaf bases); usually scapose. | Perennials; caudex simple or branched (branches short); not scapose. |
Stems | unbranched, (0.2–)0.3–1(–1.3) dm, glabrous. |
unbranched or branched distally, (0.6–)1–3(–3.8) dm, pubescent throughout, trichomes simple, 0.5–1mm, with 4–10-rayed ones, 0.05–0.2 mm (mostly branched on basal parts). |
Basal leaves | rosulate; petiole (obscure), margin ciliate, (trichomes simple or 2-rayed, 0.25–0.6 mm); blade linear to oblanceolate or narrowly obovate, (0.3–)0.4–1.2(–1.6) cm × 1–3(–4) mm, margins usually entire, rarely toothed, surfaces abaxially pubescent or glabrous, trichomes simple, sometimes with fewer, short-stalked, 2-rayed ones, (midvein prominent), adaxially often glabrous. |
rosulate; petiolate; petiole ciliate throughout; blade linear to oblanceolate or oblong, (0.5–)0.8–2(–3.5) cm × 1.5–4(–11) mm, margins entire or dentate, (base and apex ciliate, trichomes simple, 0.3–0.8 mm), surfaces pubescent with short-stalked, 4–12-rayed trichomes, 0.1–0.3 mm. |
Cauline leaves | 0–2; sessile; blade oblong to ovate, margins entire, (ciliate). |
3–10(–17), (not overlapping); sessile; blade lanceolate to ovate or oblong, margins entire or dentate, surfaces pubescent as basal (adaxially with simple and forked trichomes near blade base). |
Racemes | (2 or) 3–11(–14)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous. |
(10–)15–47(–63)-flowered, basally bracteate, often considerably elongated in fruit; rachis not flexuous, densely pubescent, trichomes 4–10-rayed (0.05–0.2 mm), and fewer simple ones. |
Flowers | sepals (green or purplish), ovate, 1.2–2.2 mm, glabrous or pubescent, (trichomes simple); petals white, spatulate, 2–2.5 × 0.8–1.5 mm; anthers ovate, 0.2–0.25 mm. |
sepals (green or lavender), oblong to ovate, 1.5–2 mm, pubescent, (trichomes simple and few-rayed); petals white, oblanceolate to spatulate, 2.3–3.7(–4.5) × 0.7–1.7 mm; anthers ovate, 0.1–0.2 mm. |
Fruiting pedicels | divaricate-ascending, often straight, (1–)2–5(–6) mm, glabrous. |
suberect or ascending, straight, 2–5(–10) mm, pubescent as rachis. |
Fruits | elliptic-lanceolate to oblong, plane, flattened, 3–8(–9) × 1.5–2 mm; valves glabrous; ovules 12–24 per ovary; style 0.05–0.2(–0.3) mm. |
(subappressed to rachis), linear-lanceolate to linear or, rarely, ovate-oblong, slightly twisted or plane, flattened, (5–)6–11 × 1.5–2(–2.5) mm; valves pubescent, trichomes short-stalked, 3–7-rayed, 0.05–0.3 mm; ovules 28–48 per ovary; style 0.1–0.6 mm. |
Seeds | oblong to elliptic, 0.8–1 × 0.5–0.6 mm. |
ovoid, 0.5–0.7(–0.9) × 0.3–0.5 mm. |
2n | = 16. |
= 32. |
Draba fladnizensis |
Draba cana |
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Phenology | Flowering Jun–Aug. | Flowering (May-)Jun–Aug. |
Habitat | Rock outcrops and talus, alpine meadows, sandy gravel | Rock outcrops and talus, open prairie benchlands, roadsides, meadows, alpine tundra |
Elevation | 0-1400 m at higher latitudes, 3000-3800 m at lower latitudes (0-4600 ft at higher latitudes, 9800-12500 ft at lower latitudes) | 0-4100 m (0-13500 ft) |
Distribution |
AK; CO; UT; WY; BC; NT; NU; QC; YT; Greenland; c Europe; s Europe; Asia; circumpolar and high alpine areas
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AK; CA; CO; ID; ME; MI; MT; NH; NM; NV; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NF; NT; ON; QC; SK; YT; Greenland
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Discussion | Rollins reduced Draba pattersonii to a variety of D. fladnizensis and separated the two primarily on plant size and minor differences in fruit shape. Examination of D. fladnizensis specimens collected throughout Europe and North America reveals that the alleged differences between the two taxa are artificial. The type material of D. pattersonii, which was collected in Colorado, is a mixture of plants highly variable in their type of indumentum. The specimens have no flowers, but the habit, fruits, and leaves are nearly indistinguishable from those of D. fladnizensis from higher latitudes. In the absence of flowers, the white-flowered Draba fladnizensis (2n = 16) is often confused with the yellow-flowered D. crassifolia (2n = 40). The latter is an annual or short-lived perennial that rarely forms a well-developed caudex, whereas D. fladnizensis almost always produces a distinct caudex. Although most individuals of both species are scapose, they occasionally produce one or two cauline leaves. The cauline leaves are usually glabrous in D. crassifolia and ciliate in D. fladnizensis; in the latter, the distalmost cauline leaf usually subtends the proximalmost flower. Finally, the seeds in D. fladnizensis are slightly larger (0.8–0.1 × 0.5–0.6 versus 0.7–0.8 × 0.4–0.5 mm) than those of D. crassifolia, though the reliability of this distinction needs to be examined in greater detail. N. H. Holmgren (2005b) reported D. fladnizensis from central Nevada, but we have not seen any material of the species from that state. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The limits of Draba cana have long been confused, and the species was treated as a synonym of the Himalayan D. lanceolata Royle (M. L. Fernald 1934; C. L. Hitchcock 1941) or as a variety of the western North American D. breweri (R. C. Rollins 1993). However, G. A. Mulligan (1971) clearly demonstrated that all three are distinct and should be maintained. Some Utah plants corresponding to the type of D. valida have shorter and wider oblong-ovate fruits. In all other aspects, they are indistinguishable from D. cana. Additional studies are needed to establish whether such plants should be formally recognized. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 307. | FNA vol. 7, p. 298. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. fladnizensis var. pattersonii, D. pattersonii, D. pattersonii var. hirticaulis, D. wahlenbergii | D. breweri var. cana, D. valida |
Name authority | Wulfen: in N. J. Jacquin, Misc. Austriac. 1: 147, plate 17, fig. 1. (1778) | Rydberg: Bull. Torrey Bot. Club 29: 241. (1902) |
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