Draba fladnizensis |
Draba arctica |
|
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arctic Draba, Austrian Draba, Austrian whitlow-grass |
arctic whitlowgrass |
|
Habit | Perennials; (sometimes cespitose); caudex simple or branched (with persistent leaf bases); usually scapose. | Perennials; (cespitose, not pulvinate); caudex simple or branched; rarely scapose. |
Stems | unbranched, (0.2–)0.3–1(–1.3) dm, glabrous. |
unbranched, 0.3–1.7(–2.4) dm, pubescent, trichomes 4–10-rayed (rays sometimes branched), 0.1–0.3 mm, with fewer, simple ones, to 0.7 mm. |
Basal leaves | rosulate; petiole (obscure), margin ciliate, (trichomes simple or 2-rayed, 0.25–0.6 mm); blade linear to oblanceolate or narrowly obovate, (0.3–)0.4–1.2(–1.6) cm × 1–3(–4) mm, margins usually entire, rarely toothed, surfaces abaxially pubescent or glabrous, trichomes simple, sometimes with fewer, short-stalked, 2-rayed ones, (midvein prominent), adaxially often glabrous. |
rosulate; petiolate; petiole base ciliate, (trichomes simple, 0.2–0.8 mm); blade oblanceolate to narrowly obovate, 0.4–2.3(–3) cm × 1.5–6.5 mm, margins usually entire, rarely with 1 tooth on each side, surfaces densely pubescent with minutely stalked, 8–12-rayed, stellate trichomes, 0.2–0.4 mm, sometimes with coarser, simple or 2-rayed ones, (midvein distinct abaxially). |
Cauline leaves | 0–2; sessile; blade oblong to ovate, margins entire, (ciliate). |
0 or 1(–3); sessile; blade ovate or oblong to lanceolate, margins entire, surfaces pubescent as basal. |
Racemes | (2 or) 3–11(–14)-flowered, usually ebracteate, rarely proximalmost flowers bracteate, elongated in fruit; rachis not flexuous, glabrous. |
3–18(–25)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals (green or purplish), ovate, 1.2–2.2 mm, glabrous or pubescent, (trichomes simple); petals white, spatulate, 2–2.5 × 0.8–1.5 mm; anthers ovate, 0.2–0.25 mm. |
sepals ovate, 2–2.5 mm, pubescent, (trichomes short-stalked, 2- or 3-rayed, and simple); petals white, spatulate to obovate, 3.5–6 × 1.8–2 mm; anthers ovate, 0.3–0.4 mm. |
Fruiting pedicels | divaricate-ascending, often straight, (1–)2–5(–6) mm, glabrous. |
divaricate-ascending or ascending, straight, (1.5–)2–6 mm, pubescent, trichomes stalked, stellate, and, sometimes, simple. |
Fruits | elliptic-lanceolate to oblong, plane, flattened, 3–8(–9) × 1.5–2 mm; valves glabrous; ovules 12–24 per ovary; style 0.05–0.2(–0.3) mm. |
oblong to lanceolate, plane, slightly flattened, (5–)6–11 × 2–3 mm; valves pubescent, trichomes short-stalked, 2–5-rayed (some rays branched), 0.05–0.3 mm; ovules 20–32 per ovary; style 0.1–1 mm (stigma distinctly wider than style). |
Seeds | oblong to elliptic, 0.8–1 × 0.5–0.6 mm. |
ovoid, 0.8–1.1 × (0.6–)0.7–0.8 mm. |
2n | = 16. |
= 80. |
Draba fladnizensis |
Draba arctica |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Jul. |
Habitat | Rock outcrops and talus, alpine meadows, sandy gravel | Clay flats, gravel, beaches, limestone outcrops, talus |
Elevation | 0-1400 m at higher latitudes, 3000-3800 m at lower latitudes (0-4600 ft at higher latitudes, 9800-12500 ft at lower latitudes) | 0-500 m (0-1600 ft) |
Distribution |
AK; CO; UT; WY; BC; NT; NU; QC; YT; Greenland; c Europe; s Europe; Asia; circumpolar and high alpine areas
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NU; Greenland; Europe (Norway [Svalbard]) |
Discussion | Rollins reduced Draba pattersonii to a variety of D. fladnizensis and separated the two primarily on plant size and minor differences in fruit shape. Examination of D. fladnizensis specimens collected throughout Europe and North America reveals that the alleged differences between the two taxa are artificial. The type material of D. pattersonii, which was collected in Colorado, is a mixture of plants highly variable in their type of indumentum. The specimens have no flowers, but the habit, fruits, and leaves are nearly indistinguishable from those of D. fladnizensis from higher latitudes. In the absence of flowers, the white-flowered Draba fladnizensis (2n = 16) is often confused with the yellow-flowered D. crassifolia (2n = 40). The latter is an annual or short-lived perennial that rarely forms a well-developed caudex, whereas D. fladnizensis almost always produces a distinct caudex. Although most individuals of both species are scapose, they occasionally produce one or two cauline leaves. The cauline leaves are usually glabrous in D. crassifolia and ciliate in D. fladnizensis; in the latter, the distalmost cauline leaf usually subtends the proximalmost flower. Finally, the seeds in D. fladnizensis are slightly larger (0.8–0.1 × 0.5–0.6 versus 0.7–0.8 × 0.4–0.5 mm) than those of D. crassifolia, though the reliability of this distinction needs to be examined in greater detail. N. H. Holmgren (2005b) reported D. fladnizensis from central Nevada, but we have not seen any material of the species from that state. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As circumscribed here, Draba arctica consists of two distinct elements often recognized as subspecies (arctica and ostenfeldii; T. W. Böcher 1966); perhaps they are better treated as distinct species (D. arctica and D. ovibovina). Lack of adequate material and inability to examine all of the types in this complex prevent us from doing so. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 307. | FNA vol. 7, p. 291. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. fladnizensis var. pattersonii, D. pattersonii, D. pattersonii var. hirticaulis, D. wahlenbergii | D. arctica subsp. ostenfeldii, D. arctica var. ostenfeldii, D. cinerea var. arctica, D. ostenfeldii, D. ostenfeldii var. ovibovina, D. ovibovina |
Name authority | Wulfen: in N. J. Jacquin, Misc. Austriac. 1: 147, plate 17, fig. 1. (1778) | J. Vahl: in G. C. Oeder et al., Fl. Dan. 13(39): plate 5, 2294. (1840) |
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