|
clawless Draba
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Heller's Draba
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| Perennials; (densely cespitose); caudex branched (covered with persistent, somewhat thickened, dry petioles); not scapose. |
Biennials or perennials; (short-lived); caudex simple or branched; not scapose. |
unbranched, 0.15–0.7 dm, glabrous throughout or sparsely pubescent, trichomes simple, 0.3–0.9 mm. |
branched, (0.1–)1.5–4.3(–5.1) dm, hirsute throughout, trichomes simple, 0.4–1.3(–1.8) mm, with short-stalked to subsessile, 3–5-rayed ones, 0.2–0.4 mm, (some slightly coarser, 2-rayed). |
rosulate; petiolate; petiole ciliate, (trichomes simple, 0.3–1.1 mm); blade linear to linear-oblanceolate, (0.5–)0.8–2(–2.5) cm × 1–3 mm, margins entire, (pubescent as petiole), surfaces abaxially sparsely pubescent with simple and stalked, 2- or 3-rayed trichomes, 0.1–0.4 mm, adaxially glabrous or subapically sparsely pubescent with simple trichomes. |
rosulate; petiolate; petiole base ciliate or not; blade oblanceolate to obovate, 0.9–4.1(–5.2) cm × 2–7(–10) mm, margins entire or dentate, surfaces pubescent, abaxially with stalked, cruciform, and fewer 3–5-rayed trichomes, 0.1–0.6 mm, adaxially with cruciform and simple trichomes, 0.4–1.3 mm, and smaller 2-rayed ones. |
1–4; sessile; blade linear to linear-oblanceolate, margins entire, surfaces pubescent as basal. |
(8–)12–31(–43); sessile; blade ovate to lanceolate or oblong, margins usually dentate, rarely subentire, surfaces pubescent as basal. |
4–13(–20)-flowered, ebracteate, elongated in fruit; rachis not flexuous, usually glabrous, rarely sparsely pubescent, trichomes simple, to 1 mm. |
10–52(–83)-flowered, usually ebracteate, rarely proximalmost 1 or 2 flowers bracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
sepals ovate, 2–2.5 mm, pubescent, (trichomes simple); petals (erect), yellow, obovate, 2–2.5(–3) × 1.5–2 mm, (not clawed); anthers oblong, 0.6–0.9 mm. |
sepals oblong, 2.5–4 mm, pubescent, (trichomes simple and stalked, 2-rayed); petals yellow, oblanceolate, 5–7 × 1.5–2.2 mm; anthers oblong, 0.5–0.8 mm. |
divaricate-ascending, straight or slightly curved upward, 1–6 mm, usually glabrous, rarely with few, simple trichomes. |
horizontal to divaricate-ascending, straight or slightly curved upward, 4–10(–13) mm, pubescent as rachis abaxially, glabrous adaxially. |
lanceolate to oblong-lanceolate, plane, flattened, 5–13 × 1.5–3 mm; valves glabrous; ovules 20–24 per ovary; style 0.3–1 mm. |
(not appressed to rachis), lanceolate to ovate or oblong-lanceolate, slightly to strongly twisted or plane, flattened, 5–15 × 2–3.5 mm; valves puberulent, trichomes simple and subsessile, 2(–4)-rayed, 0.03–0.25(–0.8) mm; ovules 14–28 per ovary; style (1–)1.5–3.5 mm. |
ovoid, 1–1.3 × 0.6–0.8 mm. |
oblong, 1–1.3 × 0.6–0.8 mm. |
= 56 ± 5. |
= 18. |
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| Flowering Jul–Aug. |
Flowering Jun–Sep. |
| Open knolls, talus, gravelly alpine slopes, rocky alpine tundra |
Oak and pine-fir woodlands, aspen groves, rocky meadows |
| 3600-4300 m (11800-14100 ft) |
2100-3600 m (6900-11800 ft) |
|
CO |
AZ; CO; NM; Mexico (Nuevo León)
|
Of conservation concern. Draba exunguiculata occupies alpine areas of Clear Creek, El Paso, Grand, and Summit counties in central Colorado. Draba grayana is found in the same general area; both species are apomicts that occasionally grow sympatrically. Draba exunguiculata is easily distinguished from D. grayana by having stems and fruiting pedicels glabrous or sparsely pubescent with simple trichomes (versus moderately to densely pubescent with simple and 2- or 3-rayed trichomes), and by having erect, non-clawed petals, 2–3 mm and slightly longer than sepals (versus flared, clawed petals 3–4.5 mm and nearly twice as long as sepals). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba helleriana is highly variable and was divided by C. L. Hitchcock (1941) into four varieties. For a discussion of those and the circumscription of the species, see I. A. Al-Shehbaz and M. D. Windham (2007). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 7, p. 306. |
FNA vol. 7, p. 311. |
| Brassicaceae > tribe Arabideae > Draba |
Brassicaceae > tribe Arabideae > Draba |
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis |
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis |
| D. chrysantha var. exunguiculata, D. chrysantha |
D. aurea var. tylosa, D. helleriana var. leiocarpa, D. helleriana var. neomexicana, D. helleriana var. patens, D. helleriana var. pinetorum, D. neomexicana, D. neomexicana var. robusta, D. pallida, D. patens, D. pinetorum, D. stylosa |
| (O. E. Schulz) C. L. Hitchcock: Revis. Drabas W. N. Amer., 46. (1941) |
Greene: Pittonia 4: 17. (1899) |
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