Flora of North America species comparison

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Draba exunguiculata

Draba grandis

clawless Draba

north Pacific Draba
Habit Perennials; (densely cespitose); caudex branched (covered with persistent, somewhat thickened, dry petioles); not scapose. Perennials; caudex simple or branched (fleshy, with persistent leaf bases); not scapose.
Stems

unbranched, 0.15–0.7 dm, glabrous throughout or sparsely pubescent, trichomes simple, 0.3–0.9 mm.

(decumbent), unbranched, (0.2–)0.5–2.7(–3.7) dm, pubescent throughout, trichomes 2–4-rayed, 0.05–0.2 mm.
Basal leaves

rosulate;

petiolate;

petiole ciliate, (trichomes simple, 0.3–1.1 mm);

blade linear to linear-oblanceolate, (0.5–)0.8–2(–2.5) cm × 1–3 mm, margins entire, (pubescent as petiole), surfaces abaxially sparsely pubescent with simple and stalked, 2- or 3-rayed trichomes, 0.1–0.4 mm, adaxially glabrous or subapically sparsely pubescent with simple trichomes.

rosulate; long-petiolate;

petiole (winged, (1–)4–15 cm), often not ciliate (or ciliate to blade apex, trichomes simple, 0.3–0.8 mm);

blade (somewhat fleshy), oblanceolate to spatulate or obovate, (1–)2–11(–17) cm × (5–)8–30(–45) mm, margins often coarsely dentate, (pubescent as petiole), surfaces usually pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.05–0.25 mm, adaxially similar, or also with simple and long-stalked, 2-rayed trichomes, to 0.8 mm, rarely glabrescent, with mostly simple and 2-rayed trichomes.
Cauline leaves

1–4;

sessile;

blade linear to linear-oblanceolate, margins entire, surfaces pubescent as basal.

2–12(–16);

sessile or petiolate;

blade oblanceolate to obovate, margins dentate or entire, surfaces pubescent as basal.
Racemes

4–13(–20)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, usually glabrous, rarely sparsely pubescent, trichomes simple, to 1 mm.

5–26(–32)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.
Flowers

sepals ovate, 2–2.5 mm, pubescent, (trichomes simple);

petals (erect), yellow, obovate, 2–2.5(–3) × 1.5–2 mm, (not clawed);

anthers oblong, 0.6–0.9 mm.

sepals broadly ovate, 3–4 mm, pubescent, (trichomes simple and short-stalked, 2-rayed);

petals yellow, oblanceolate to spatulate, 4.5–7 × 1.8–3 mm;

anthers oblong, 0.7–1 mm.
Fruiting pedicels

divaricate-ascending, straight or slightly curved upward, 1–6 mm, usually glabrous, rarely with few, simple trichomes.

horizontal to divaricate-ascending or ascending, usually straight, rarely curved upward, (5–)10–22(–27) mm, pubescent, trichomes 2–4-rayed (0.05–0.3 mm), sometimes with simple and spurred ones.
Fruits

lanceolate to oblong-lanceolate, plane, flattened, 5–13 × 1.5–3 mm;

valves glabrous;

ovules 20–24 per ovary;

style 0.3–1 mm.

oblong to lanceolate, or ovate to suborbicular, slightly twisted or plane, flattened, (6–)10–20(–25) × 4–7(–9) mm;

valves glabrous;

ovules 24–52 per ovary;

style (0.2–)0.4–1.6(–2) mm.
Seeds

ovoid, 1–1.3 × 0.6–0.8 mm.

ovoid, 1.4–2 × 0.8–1.2 mm.
2n

= 56 ± 5.

= 36.

Draba exunguiculata

Draba grandis
Phenology Flowering Jul–Aug. Flowering May–Jul.
Habitat Open knolls, talus, gravelly alpine slopes, rocky alpine tundra Rocky bluffs above salt-water beaches, loamy seaside banks, sea bird rookeries, coastal herbaceous tundra and sandy blowouts
Elevation 3600-4300 m (11800-14100 ft) 0-50(-200) m (0-200(-700) ft)
Distribution
from FNA
CO
[BONAP county map]
from FNA
AK; BC; e Asia (Kuril and Ratmanov islands, Russian Far East)
[BONAP county map]
Discussion

Of conservation concern.

Draba exunguiculata occupies alpine areas of Clear Creek, El Paso, Grand, and Summit counties in central Colorado. Draba grayana is found in the same general area; both species are apomicts that occasionally grow sympatrically. Draba exunguiculata is easily distinguished from D. grayana by having stems and fruiting pedicels glabrous or sparsely pubescent with simple trichomes (versus moderately to densely pubescent with simple and 2- or 3-rayed trichomes), and by having erect, non-clawed petals, 2–3 mm and slightly longer than sepals (versus flared, clawed petals 3–4.5 mm and nearly twice as long as sepals).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Almost all recent North American authors have used the name Draba hyperborea for this species. A. N. Berkutenko (1995) clearly showed that the type of that name belongs to an entirely different species that she placed in the genus Schivereckia Andrzejowski ex de Candolle. Draba grandis thus becomes the correct name for the North American taxon. Except for its fleshy leaves, Nesodraba is indistinguishable morphologically and molecularly from other species of Draba.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 7, p. 306. FNA vol. 7, p. 309.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. chrysantha var. exunguiculata, D. chrysantha Cochlearia siliquosa, Cochlearia spathulata, D. greenei, D. hatchiae, D. hyperborea var. spathulata, D. spathulata, Nesodraba grandis, Nesodraba megalocarpa, Nesodraba siliquosa
Name authority (O. E. Schulz) C. L. Hitchcock: Revis. Drabas W. N. Amer., 46. (1941) Langsdorff ex de Candolle: Syst. Nat. 2: 355. (1821)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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