Draba exunguiculata |
Draba cana |
|
---|---|---|
clawless Draba |
Brewer's Draba, canescent Draba, canescent whitlow-mustard, cushion Draba, hoary Draba, hoary whitlow-grass, lance-leaf Draba, lance-leaf whitlow-grass |
|
Habit | Perennials; (densely cespitose); caudex branched (covered with persistent, somewhat thickened, dry petioles); not scapose. | Perennials; caudex simple or branched (branches short); not scapose. |
Stems | unbranched, 0.15–0.7 dm, glabrous throughout or sparsely pubescent, trichomes simple, 0.3–0.9 mm. |
unbranched or branched distally, (0.6–)1–3(–3.8) dm, pubescent throughout, trichomes simple, 0.5–1mm, with 4–10-rayed ones, 0.05–0.2 mm (mostly branched on basal parts). |
Basal leaves | rosulate; petiolate; petiole ciliate, (trichomes simple, 0.3–1.1 mm); blade linear to linear-oblanceolate, (0.5–)0.8–2(–2.5) cm × 1–3 mm, margins entire, (pubescent as petiole), surfaces abaxially sparsely pubescent with simple and stalked, 2- or 3-rayed trichomes, 0.1–0.4 mm, adaxially glabrous or subapically sparsely pubescent with simple trichomes. |
rosulate; petiolate; petiole ciliate throughout; blade linear to oblanceolate or oblong, (0.5–)0.8–2(–3.5) cm × 1.5–4(–11) mm, margins entire or dentate, (base and apex ciliate, trichomes simple, 0.3–0.8 mm), surfaces pubescent with short-stalked, 4–12-rayed trichomes, 0.1–0.3 mm. |
Cauline leaves | 1–4; sessile; blade linear to linear-oblanceolate, margins entire, surfaces pubescent as basal. |
3–10(–17), (not overlapping); sessile; blade lanceolate to ovate or oblong, margins entire or dentate, surfaces pubescent as basal (adaxially with simple and forked trichomes near blade base). |
Racemes | 4–13(–20)-flowered, ebracteate, elongated in fruit; rachis not flexuous, usually glabrous, rarely sparsely pubescent, trichomes simple, to 1 mm. |
(10–)15–47(–63)-flowered, basally bracteate, often considerably elongated in fruit; rachis not flexuous, densely pubescent, trichomes 4–10-rayed (0.05–0.2 mm), and fewer simple ones. |
Flowers | sepals ovate, 2–2.5 mm, pubescent, (trichomes simple); petals (erect), yellow, obovate, 2–2.5(–3) × 1.5–2 mm, (not clawed); anthers oblong, 0.6–0.9 mm. |
sepals (green or lavender), oblong to ovate, 1.5–2 mm, pubescent, (trichomes simple and few-rayed); petals white, oblanceolate to spatulate, 2.3–3.7(–4.5) × 0.7–1.7 mm; anthers ovate, 0.1–0.2 mm. |
Fruiting pedicels | divaricate-ascending, straight or slightly curved upward, 1–6 mm, usually glabrous, rarely with few, simple trichomes. |
suberect or ascending, straight, 2–5(–10) mm, pubescent as rachis. |
Fruits | lanceolate to oblong-lanceolate, plane, flattened, 5–13 × 1.5–3 mm; valves glabrous; ovules 20–24 per ovary; style 0.3–1 mm. |
(subappressed to rachis), linear-lanceolate to linear or, rarely, ovate-oblong, slightly twisted or plane, flattened, (5–)6–11 × 1.5–2(–2.5) mm; valves pubescent, trichomes short-stalked, 3–7-rayed, 0.05–0.3 mm; ovules 28–48 per ovary; style 0.1–0.6 mm. |
Seeds | ovoid, 1–1.3 × 0.6–0.8 mm. |
ovoid, 0.5–0.7(–0.9) × 0.3–0.5 mm. |
2n | = 56 ± 5. |
= 32. |
Draba exunguiculata |
Draba cana |
|
Phenology | Flowering Jul–Aug. | Flowering (May-)Jun–Aug. |
Habitat | Open knolls, talus, gravelly alpine slopes, rocky alpine tundra | Rock outcrops and talus, open prairie benchlands, roadsides, meadows, alpine tundra |
Elevation | 3600-4300 m (11800-14100 ft) | 0-4100 m (0-13500 ft) |
Distribution |
CO |
AK; CA; CO; ID; ME; MI; MT; NH; NM; NV; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NF; NT; ON; QC; SK; YT; Greenland
|
Discussion | Of conservation concern. Draba exunguiculata occupies alpine areas of Clear Creek, El Paso, Grand, and Summit counties in central Colorado. Draba grayana is found in the same general area; both species are apomicts that occasionally grow sympatrically. Draba exunguiculata is easily distinguished from D. grayana by having stems and fruiting pedicels glabrous or sparsely pubescent with simple trichomes (versus moderately to densely pubescent with simple and 2- or 3-rayed trichomes), and by having erect, non-clawed petals, 2–3 mm and slightly longer than sepals (versus flared, clawed petals 3–4.5 mm and nearly twice as long as sepals). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The limits of Draba cana have long been confused, and the species was treated as a synonym of the Himalayan D. lanceolata Royle (M. L. Fernald 1934; C. L. Hitchcock 1941) or as a variety of the western North American D. breweri (R. C. Rollins 1993). However, G. A. Mulligan (1971) clearly demonstrated that all three are distinct and should be maintained. Some Utah plants corresponding to the type of D. valida have shorter and wider oblong-ovate fruits. In all other aspects, they are indistinguishable from D. cana. Additional studies are needed to establish whether such plants should be formally recognized. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 306. | FNA vol. 7, p. 298. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. chrysantha var. exunguiculata, D. chrysantha | D. breweri var. cana, D. valida |
Name authority | (O. E. Schulz) C. L. Hitchcock: Revis. Drabas W. N. Amer., 46. (1941) | Rydberg: Bull. Torrey Bot. Club 29: 241. (1902) |
Web links |
|