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bitter root Draba

Standley's Draba

Habit Perennials; (densely pulvinate); caudex branched (branches elongated, loose, with persistent leaf remains, terminating in flowering or sterile shoots); scapose. Perennials; (densely pulvinate); caudex branched (with persistent, thickened petioles, branches compact); not scapose.
Stems

unbranched, (0.05–)0.2–0.6 dm, glabrous.

unbranched, 0.3–1.3(–1.7) dm, usually glabrous throughout or sparsely pubescent proximally, rarely sparsely pubescent distally, trichomes simple and 2-rayed, 0.1–0.7 mm.

Basal leaves

(densely imbricate); rosulate;

petiolate;

petiole ciliate throughout;

blade (fleshy), oblong to obovate or oblanceolate, 0.3–0.7(–1) cm × 1–2(–2.5) mm, margins entire, (ciliate, trichomes simple, 0.1–0.5 mm, apex obtuse), surfaces glabrous (midvein obscure abaxially).

(not imbricate); rosulate;

petiolate;

petiole ciliate, (trichomes usually simple, rarely 2-rayed, not setiform);

blade narrowly oblanceolate to linear-lanceolate, (strongly differentiated into blade and petiole), (1.2–)1.8–6(–8.5) cm × 1–5(–7) mm, margins entire or sparsely denticulate, (ciliate as petiole; midvein not prominent), surfaces glabrous or pubescent, usually with simple trichomes 0.1–0.8 mm, rarely 2-rayed.

Cauline leaves

0.

1–8;

sessile;

blade lanceolate to narrowly oblong, margins usually entire, surfaces pubescent as basal.

Racemes

2–8(–10)-flowered, ebracteate, (subcorymbose), slightly elongated in fruit;

rachis not flexuous, glabrous.

5–17(–23)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, usually glabrous, rarely pubescent, trichomes simple and 2-rayed, (non-crisped).

Flowers

sepals oblong, 1.5–2.2 mm, glabrous;

petals pale to bright yellow, spatulate, 3.5–4 × 1–2 mm;

anthers ovate, 0.3–0.4 mm.

sepals ovate, 2–2.5 mm, pubescent, (trichomes simple);

petals yellow, oblanceolate, 4–6 × 1.5–2 mm;

anthers oblong, 0.6–0.8 mm.

Fruiting pedicels

divaricate-ascending (not decurrent basally), straight, 4–10 mm, glabrous.

divaricate-ascending or ascending, straight, 3–9(–13) mm, usually glabrous, rarely sparsely pubescent, trichomes simple.

Fruits

ovate to oblong-elliptic, plane, flattened, 4–8 × 2–4 mm;

valves (obscurely veined), glabrous;

ovules 6–14 per ovary;

style 0.1–0.5 mm.

linear-elliptic to elliptic, twisted or plane, flattened, 5–10(–13) × 1.5–2.5 mm;

valves usually glabrous, rarely puberulent, trichomes simple, 0.05–0.1 mm;

ovules 12–24 per ovary;

style 0.7–1.4(–1.8) mm.

Seeds

ovoid, 1.2–1.5 × 0.8–1 mm.

ovoid, 1–1.2 × 0.7–0.8 mm.

Draba daviesiae

Draba standleyi

Phenology Flowering Jul–Aug. Flowering Jun–Aug.
Habitat Talus slopes, rock crevices and cracks, rocky ridges and slides, alpine meadows Igneous rock outcrops, stabilized talus slopes
Elevation 2700-2900 m (8900-9500 ft) 1800-3100 m (5900-10200 ft)
Distribution
from FNA
MT
[BONAP county map]
from FNA
AZ; NM; TX
[BONAP county map]
Discussion

Although originally described as a variety of Draba apiculata (= D. globosa), D. daviesiae is distinct morphologically. It is easily distinguished from the former by its densely pulvinate habit, obtuse leaf blades, and obscurely veined fruit valves. By contrast, D. globosa exhibits a cespitose but non-pulvinate habit, acute leaf blades, and prominently veined fruit valves. Draba daviesiae is known from the Bitterroot Mountains in Ravalli County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Draba standleyi is a distinctive species that is sporadically distributed in the mountains of southwestern United States. It is known from the Chiricahua Mountains (Cochise County, southeastern Arizona), the Organ Mountains and Black Range (Dona Ana and Sierra counties, south-central New Mexico), and the Davis Mountains (Jeff Davis County, western Texas). It has not been reported from Mexico, though it is very likely to occur there.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 305. FNA vol. 7, p. 340.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. apiculata var. daviesiae, D. densifolia var. daviesiae D. gilgiana, D. chrysantha var. gilgiana
Name authority (C. L. Hitchcock) Rollins: Contr. Gray Herb. 214: 5. (1984) J. F. Macbride & Payson: Ann. Missouri Bot. Gard. 5: 150. (1918)
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