Draba cuneifolia var. integrifolia |
Draba cuneifolia |
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wedgeleaf Draba |
wedge leaf Draba |
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Habit | Annuals; scapose or subscapose. | |||||||||
Stems | pubescent with 3- or 4-rayed trichomes. |
(simple to many from or near base), unbranched, (0.2–) 0.3–2.7(–3.7) dm, hirsute or pubescent throughout, trichomes 3–4-rayed, 0.05–0.4 mm, (sometimes mixed proximally with simple or spurred ones, 0.5–1.2 mm). |
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Basal leaves | not rosulate; petiole (obscure), not ciliate; blade oblanceolate to spatulate or broadly obovate, (0.4–)1–3.5(–5) cm × (2–)6–20(–28) mm, margins dentate (in distal 1/2), surfaces pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.1–0.7 mm, adaxially similar or also with fewer, simple trichomes, 0.4–0.7 mm. |
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Cauline leaves | 0–6 (on proximal 1/3 of stem); blade similar to basal. |
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Racemes | to 3/4 of scape. |
10–50(–70)-flowered (throughout or on distal 1/3 of scape), ebracteate, elongated in fruit; rachis not flexuous, densely pubescent, trichomes 2–4-rayed. |
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Flowers | (late ones cleistogamous, apetalous); sepals (green or pink), oblong, 1.5–2.5 mm, glabrous or pubescent, (trichomes simple); petals white, spatulate, (2–)2.5–4.5(–5) × 1–2 mm, (emarginate or obtuse); anthers ovate to oblong, (0.1–)0.25–0.4 mm. |
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Fruiting pedicels | horizontal to divaricate-ascending, straight, (1–)2–7(–10) mm, pubescent as rachis. |
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Fruits | oblong to linear or lanceolate, (5–)7–12 mm; valves: trichomes stalked and 4-rayed, sometimes with 2- or 3-rayed ones; ovules 20–44 per ovary. |
oblong to linear or lanceolate to broadly ovate, plane, flattened, (3–)6–12(–16) × 1.7–2.7(–3) mm; valves usually puberulent, rarely glabrous, trichomes simple, antrorse, 0.1–0.3 mm, (rarely with 2-rayed ones, or all trichomes short-stalked, 4-rayed, cruciform); ovules (12–)24–66(–72) per ovary; style 0.01–0.3(–0.4) mm. |
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Seeds | broadly ovoid, 0.5–0.7 × 0.4–0.5 mm. |
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2n | = 30, 32. |
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Draba cuneifolia var. integrifolia |
Draba cuneifolia |
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Phenology | Flowering Jan–Mar. | |||||||||
Habitat | Rocky slopes and gravelly soil in desert scrub communities | |||||||||
Elevation | 200-1800 m (700-5900 ft) | |||||||||
Distribution |
AZ; CA; NV; TX; UT; Mexico (Baja California, Nuevo León, Sonora) |
AL; AR; AZ; CA; CO; FL; IL; KS; KY; LA; MO; MS; NC; NE; NM; NV; OH; OK; PA; SD; TN; TX; UT; n Mexico |
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Discussion | Varieties 3 (3 in the flora). Draba cuneifolia is highly variable in fruit shape and size, style length, ovule number per ovary, type of fruit indumentum, and length of the fruiting raceme in relation to the rest of the scape. Reported chromosome numbers also vary, but it is currently unclear whether this is the result of biologically relevant processes or erroneous counts. R. L. Hartman et al. (1975) divided the species into three varieties, a position followed by subsequent authors (e.g., R. C. Rollins 1993). Typical var. sonorae is rather distinct (see key below) and may deserve recognition as a separate species, as by O. E. Schulz (1927). Critical, population-based, molecular studies are needed to assess this. In the absence of such studies, we tentatively follow Hartman et al., recognizing three varieties that are distinguished primarily by the types of trichomes found on the fruits. Glabrous-fruited forms occasionally encountered in all three varieties can be difficult to identify. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 304. | FNA vol. 7, p. 303. | ||||||||
Parent taxa | ||||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | D. integrifolia, D. sonorae var. integrifolia | |||||||||
Name authority | S. Watson: Proc. Amer. Acad. Arts 23: 256. (1888) | Nuttall ex Torrey & A. Gray: Fl. N. Amer. 1: 108. (1838) | ||||||||
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