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wedgeleaf Draba

wedge leaf Draba

Habit Annuals; scapose or subscapose.
Stems

pubescent with 3- or 4-rayed trichomes.

(simple to many from or near base), unbranched, (0.2–) 0.3–2.7(–3.7) dm, hirsute or pubescent throughout, trichomes 3–4-rayed, 0.05–0.4 mm, (sometimes mixed proximally with simple or spurred ones, 0.5–1.2 mm).

Basal leaves

not rosulate;

petiole (obscure), not ciliate;

blade oblanceolate to spatulate or broadly obovate, (0.4–)1–3.5(–5) cm × (2–)6–20(–28) mm, margins dentate (in distal 1/2), surfaces pubescent, abaxially with stalked, 2–4-rayed trichomes, 0.1–0.7 mm, adaxially similar or also with fewer, simple trichomes, 0.4–0.7 mm.

Cauline leaves

0–6 (on proximal 1/3 of stem);

blade similar to basal.

Racemes

to 3/4 of scape.

10–50(–70)-flowered (throughout or on distal 1/3 of scape), ebracteate, elongated in fruit;

rachis not flexuous, densely pubescent, trichomes 2–4-rayed.

Flowers

(late ones cleistogamous, apetalous);

sepals (green or pink), oblong, 1.5–2.5 mm, glabrous or pubescent, (trichomes simple);

petals white, spatulate, (2–)2.5–4.5(–5) × 1–2 mm, (emarginate or obtuse);

anthers ovate to oblong, (0.1–)0.25–0.4 mm.

Fruiting pedicels

horizontal to divaricate-ascending, straight, (1–)2–7(–10) mm, pubescent as rachis.

Fruits

oblong to linear or lanceolate, (5–)7–12 mm;

valves: trichomes stalked and 4-rayed, sometimes with 2- or 3-rayed ones;

ovules 20–44 per ovary.

oblong to linear or lanceolate to broadly ovate, plane, flattened, (3–)6–12(–16) × 1.7–2.7(–3) mm;

valves usually puberulent, rarely glabrous, trichomes simple, antrorse, 0.1–0.3 mm, (rarely with 2-rayed ones, or all trichomes short-stalked, 4-rayed, cruciform);

ovules (12–)24–66(–72) per ovary;

style 0.01–0.3(–0.4) mm.

Seeds

broadly ovoid, 0.5–0.7 × 0.4–0.5 mm.

2n

= 30, 32.

Draba cuneifolia var. integrifolia

Draba cuneifolia

Phenology Flowering Jan–Mar.
Habitat Rocky slopes and gravelly soil in desert scrub communities
Elevation 200-1800 m (700-5900 ft)
Distribution
from FNA
AZ; CA; NV; TX; UT; Mexico (Baja California, Nuevo León, Sonora)
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; FL; IL; KS; KY; LA; MO; MS; NC; NE; NM; NV; OH; OK; PA; SD; TN; TX; UT; n Mexico
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Draba cuneifolia is highly variable in fruit shape and size, style length, ovule number per ovary, type of fruit indumentum, and length of the fruiting raceme in relation to the rest of the scape. Reported chromosome numbers also vary, but it is currently unclear whether this is the result of biologically relevant processes or erroneous counts. R. L. Hartman et al. (1975) divided the species into three varieties, a position followed by subsequent authors (e.g., R. C. Rollins 1993). Typical var. sonorae is rather distinct (see key below) and may deserve recognition as a separate species, as by O. E. Schulz (1927). Critical, population-based, molecular studies are needed to assess this. In the absence of such studies, we tentatively follow Hartman et al., recognizing three varieties that are distinguished primarily by the types of trichomes found on the fruits. Glabrous-fruited forms occasionally encountered in all three varieties can be difficult to identify.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit valves: trichomes usually simple, sometimes with 2-rayed ones; stems hirsute with at least some simple trichomes.
var. cuneifolia
1. Fruit valves: trichomes stalked and 4-rayed, sometimes with 2- or 3-rayed ones; stems pubescent with 3- or 4-rayed trichomes
→ 2
2. Fruits (5-)7-12 mm; ovules 20-44 per ovary; racemes to 3/4 of scape.
var. integrifolia
2. Fruits 3-6(-8) mm; ovules 12-24 per ovary; racemes on all or most of scape.
var. sonorae
Source FNA vol. 7, p. 304. FNA vol. 7, p. 303.
Parent taxa Brassicaceae > tribe Arabideae > Draba > Draba cuneifolia Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. cuneifolia var. cuneifolia, D. cuneifolia var. sonorae
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Subordinate taxa
D. cuneifolia var. cuneifolia, D. cuneifolia var. integrifolia, D. cuneifolia var. sonorae
Synonyms D. integrifolia, D. sonorae var. integrifolia
Name authority S. Watson: Proc. Amer. Acad. Arts 23: 256. (1888) Nuttall ex Torrey & A. Gray: Fl. N. Amer. 1: 108. (1838)
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