Draba crassifolia |
Draba paysonii |
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Rocky Mountain Draba, Rocky Mountain whitlow-grass, snowbed Draba, snowbed whitlow-grass, thick-leaf Draba |
Payson's Draba |
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Habit | Annuals or perennials; (short-lived); caudex branched (when present); usually scapose. | Perennials; (cespitose, densely pulvinate); caudex branched (covered with persistent leaves, branches creeping, sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched or, rarely, branched distally, (0.1–)0.3–1.1(–1.5) dm, usually glabrous throughout, rarely pubescent proximally, trichomes simple, 0.3–0.7 mm. |
unbranched, 0.05–0.3 dm, densely pubescent throughout, trichomes simple, 0.4–1.3 mm, and stalked, 2–5-rayed, stellate, 0.1–0.6 mm. |
Basal leaves | rosulate; petiolate; petiole ciliate throughout; blade oblanceolate to obovate, (0.2–) 0.5–2.5(–3) cm × (1–)2–4(–6) mm, margins usually entire, rarely denticulate, (sometimes ciliate), surfaces glabrous or sparsely pubescent, with simple and 2-rayed trichomes, 0.3–0.9 mm. |
(imbricate); rosulate; sessile; blade linear to linear-oblanceolate, 0.4–1.2 cm × 0.8–1.5 mm, margins entire, (ciliate, trichomes simple and spurred, (0.4–)0.6–1.4 mm), surfaces pubescent, abaxially (dense) with stalked, 2–12-rayed trichomes, 0.1–0.8 mm, adaxially with simple and 2-rayed ones, 0.3–1 mm, (midvein prominent). |
Cauline leaves | usually 0, rarely 1; sessile; blade oblong to ovate, margins entire, surfaces glabrous. |
0. |
Racemes | (2–)4–15(–25)-flowered, ebracteate, elongated in fruit; rachis slightly flexuous or straight, glabrous. |
2–7-flowered, ebracteate, slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | (chasmogamous, petaliferous); sepals (green or purplish), ovate, 1–2 mm, glabrous; petals yellow (often fading white), oblanceolate, 1.5–2.5(–3) × 0.5–1 mm; anthers ovate, 0.15–0.25 mm. |
sepals oblong, 2.8–3.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals pale yellow, oblanceolate to spatulate, (4–)5–6 × (1.5–)2–3 mm; anthers oblong, 0.5–0.7 mm. |
Fruiting pedicels | horizontal to divaricate-ascending, usually straight, rarely curved upward, 3–8(–10) mm (subequaling or shorter than fruit), glabrous. |
divaricate-ascending, straight, 3–9 mm, pubescent, trichomes simple (0.4–1.4 mm), and stalked, 2–5-rayed (0.1–0.6 mm). |
Fruits | usually narrowly elliptic to lanceolate, rarely linear-lanceolate, plane, flattened, (3–)5–10 × 1.5–2.5 mm; valves glabrous; ovules (8–)16–24(–30) per ovary; style 0.02–0.1 mm. |
often broadly ovoid to lanceolate, plane, slightly inflated basally, (5–)6–9 × (3–)3.7–5 mm; valves densely pubescent, trichomes simple and 2-rayed, (0.2–)0.4–1 mm, (occasionally with fewer, smaller, 4- or 5-rayed ones); ovules 8–12 per ovary; style (0.6–)0.8–1.2 mm. |
Seeds | elliptic, 0.7–0.8 × 0.4–0.5 mm. |
oblong, 1.7–2.2 × 1–1.4 mm. |
2n | = 40. |
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Draba crassifolia |
Draba paysonii |
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Phenology | Flowering Jun–Sep. | Flowering Jun–Jul. |
Habitat | Rock outcrops and talus, subalpine meadows, alpine summits and tundra, bare snow-melt areas | Limestone rock outcrops, talus, gravelly calcareous soil |
Elevation | (50-)1000-4300 m ((200-)3300-14100 ft) | 1800-3500 m (5900-11500 ft) |
Distribution |
AK; AZ; CO; ID; MT; NM; NV; UT; WA; WY; AB; BC; LB; NT; NU; QC; YT; Greenland; n Europe (Norway, Sweden)
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MT; WY |
Discussion | M. D. Windham (2004) presented morphological and chromosomal data suggesting that Draba crassifolia is an allopolyploid produced by hybridization between D. albertina and D. fladnizensis. Although the species is distinctive in large part, some individuals can be difficult to place and there is evidence of rare backcrossing (Windham, unpubl.). The attribution to Arizona is based on Schaack 345 (US) and Kearney & Peebles 12156 (US), both collected on the San Francisco Peaks in Coconino County. Draba crassifolia is found at elevations as low as 50 m in Greenland and the islands of Nunavut. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba paysonii is known from southwestern Montana (Gallatin, Glacier, Madison, and Meagher counties) and northwestern Wyoming (Fremont and Park counties). Reports of its occurrence in Alberta (e.g., G. A. Mulligan 1971b) are based on plants of D. novolympica, a species formerly treated as D. paysonii var. treleasei. Features distinguishing these two species are discussed under 70. D. novolympica. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 302. | FNA vol. 7, p. 327. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. crassifolia var. parryi, D. parryi | D. vestita |
Name authority | Graham: Edinburgh New Philos. J. 7: 182. (1829) | J. F. Macbride: Contr. Gray Herb. 56: 52. (1918) |
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