Draba corymbosa |
Draba zionensis |
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Baffin bay Draba, Baffin bay whitlow-grass, flat-top Draba |
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Habit | Perennials; (cespitose); caudex branched (with persistent leaves or leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (cespitose); caudex branched (with persistent leaf bases, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, (0.05–)0.2–0.8(–1.5) dm, pubescent throughout, trichomes simple and 2-rayed, 0.4–1 mm, with 3–5-rayed ones, 0.05–0.4 mm, (sometimes trichomes mostly simple). |
unbranched, 0.5–1.5(–1.8) dm, usually sparsely pubescent proximally, usually glabrous distally, rarely throughout, trichomes simple, 0.3–0.8 mm, often with smaller, 2–4-rayed ones. |
Basal leaves | rosulate; petiolate; petiole base and margin ciliate, (trichomes often course, simple, 0.4–1.3 mm); blade oblanceolate to obovate, 0.6–1.8 cm × 1.5–5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–6-rayed trichomes, 0.2–0.6 mm, adaxially with primarily simple and stalked, 2-rayed trichomes, to 1.1 mm, with 3–5-rayed ones, 0.2–0.4 mm. |
rosulate; shortly petiolate; petiole base sometimes ciliate (margin not ciliate, trichomes simple and 2-rayed, 0.2–0.5 mm); blade spatulate to obovate, 0.7–3.5 cm × 3–10(–12) mm, margins usually entire, rarely obscurely dentate (near apex), surfaces sparsely pubescent with stalked, (2–)4 (or 5)-rayed trichomes, 0.05–0.5 mm, (midvein obscure). |
Cauline leaves | 0. |
0. |
Racemes | 2–9(–12)-flowered, ebracteate, (corymbose), slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
14–36-flowered, ebracteate, elongated in fruit; rachis not flexuous, glabrous. |
Flowers | sepals (grayish green), broadly ovate, 2.2–3 mm, pubescent, (trichomes simple, to 1 mm, sometimes with stalked, smaller, 2–4-rayed ones); petals (broadly patent), yellow, obovate, 4–6 × 3–5 mm; anthers ovate, 0.3–0.4 mm. |
sepals ovate, 2–3 mm, glabrous or sparsely pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals orange-yellow (fading pale yellow), broadly spatulate to subovate, 5–6 × 1.8–2.8 mm; anthers ovate, 0.4–0.5 mm. |
Fruiting pedicels | divaricate-ascending, straight or slightly curved upward, 4–11(–16) mm, pubescent, trichomes simple and 2–4-rayed. |
ascending to divaricate-ascending, straight, (not expanded basally), (4–)5–12(–15) mm, glabrous. |
Fruits | oblong or ovate, plane, flattened, 6–12 × 3.5–5.5 mm; valves pubescent or puberulent, trichomes simple, 0.1–0.4 mm, (sometimes with short-stalked, 2- or 3-rayed ones); ovules 12–24 per ovary; style 0.6–1 mm (stigma distinctly wider than style). |
lanceolate to linear-lanceolate or ovate, plane (not curved), flattened, 6–12(–17) × 2–3 mm; valves glabrous; ovules 12–20 per ovary; style 0.4–1 mm. |
Seeds | (brown), ovoid, 1–1.3 × 0.6–0.9 mm. |
oblong, 1–1.3 × 0.6–0.8 mm. |
2n | = 128, 144. |
= 26. |
Draba corymbosa |
Draba zionensis |
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Phenology | Flowering Jun–Aug. | Flowering Mar–May. |
Habitat | Moist tundra, among calcareous or dolomitic rocks, gravel beaches, silt and clay terraces | Sandstone (rarely limestone) rock outcrops and sandy slopes in pinyon-juniper or pine communities |
Elevation | 0-1700 m (0-5600 ft) | 1000-2500 m (3300-8200 ft) |
Distribution |
AK; BC; NT; NU; YT; Greenland; Europe (Norway, n Russia); e Asia (Russian Far East, Siberia) |
UT |
Discussion | Draba corymbosa, which is 16-ploid or 18-ploid with x = 8, is an extremely variable species of polyphyletic, allopolyploid origin. Most individuals appear to have decaploid D. alpina (2n = 80) in their parentage, but the other genomes are provided by hexaploid (2n = 48) and octoploid (2n = 64) species (C. Brochmann et al. 1993). O. E. Schulz reduced D. corymbosa to a variety of D. alpina; R. C. Rollins (1993) treated it as a distinct species. Draba corymbosa is distinguished from D. alpina by having pubescent or puberulent (versus glabrous or glabrescent) fruits, corymbose (versus usually elongated) fruiting racemes, and abaxial leaf blade surfaces always lacking (versus usually with some) simple trichomes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
R. C. Rollins (1993) treated Draba zionensis as a variety of D. asprella, but its true relationships appear to lie with two other southern Utah endemics, D. sobolifera and D. subalpina. Draba zionensis is easily distinguished from D. subalpina by having orange-yellow (versus white) petals, and from D. asprella and D. sobolifera by its glabrous (versus pubescent) pedicels and stems distally. Nearly all populations of the species are found in and around Zion National Park in southwestern Utah (Iron, Kane, and Washington counties). A specimen supposedly from the Deep Creek Mountains (Juab County) may be mislabeled. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 301. | FNA vol. 7, p. 346. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. alpina var. bellii, D. alpina var. corymbosa, D. barbata, D. bellii, D. kjellmanii, D. macrocarpa, D. vestita | D. asprella var. zionensis |
Name authority | R. Brown ex de Candolle: Syst. Nat. 2: 343. (1821) | C. L. Hitchcock: Revis. Drabas W. N. Amer., 49, plate 2, fig. 16. (1941) |
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