Draba corymbosa |
Draba porsildii |
|
---|---|---|
Baffin bay Draba, Baffin bay whitlow-grass, flat-top Draba |
Porsild's Draba, Porsild's whitlow-grass |
|
Habit | Perennials; (cespitose); caudex branched (with persistent leaves or leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (cespitose); caudex branched (with persistent leaf remains); often scapose. |
Stems | unbranched, (0.05–)0.2–0.8(–1.5) dm, pubescent throughout, trichomes simple and 2-rayed, 0.4–1 mm, with 3–5-rayed ones, 0.05–0.4 mm, (sometimes trichomes mostly simple). |
unbranched, 0.2–1.2 dm, sparsely pubescent proximally, trichomes short-stalked, 3–5- rayed, (non-crisped), 0.2–0.4 mm, glabrous distally. |
Basal leaves | rosulate; petiolate; petiole base and margin ciliate, (trichomes often course, simple, 0.4–1.3 mm); blade oblanceolate to obovate, 0.6–1.8 cm × 1.5–5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–6-rayed trichomes, 0.2–0.6 mm, adaxially with primarily simple and stalked, 2-rayed trichomes, to 1.1 mm, with 3–5-rayed ones, 0.2–0.4 mm. |
rosulate; subsessile or shortly petiolate; petiole base and margin ciliate proximally, (trichomes simple, subsetiform, stiff, 0.3–0.9 mm); blade oblanceolate to obovate, 0.4–1.5 cm × 1–3 mm, margins entire, surfaces pubescent, abaxially with stalked, 6–12-rayed, (non-crisped) trichomes, 0.2–0.5 mm, (midvein obscure), adaxially similar, or with simple trichomes on proximal 1/2. |
Cauline leaves | 0. |
0 or 1; sessile; blade oblong to lanceolate, margins entire, surfaces pubescent as basal. |
Racemes | 2–9(–12)-flowered, ebracteate, (corymbose), slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
3–13-flowered, ebracteate, elongated in fruit; rachis not flexuous, glabrous. |
Flowers | sepals (grayish green), broadly ovate, 2.2–3 mm, pubescent, (trichomes simple, to 1 mm, sometimes with stalked, smaller, 2–4-rayed ones); petals (broadly patent), yellow, obovate, 4–6 × 3–5 mm; anthers ovate, 0.3–0.4 mm. |
sepals ovate, 1.5–2 mm, pubescent, (trichomes simple and 2-rayed); petals white, spatulate, 2–4 × 1–1.7 mm; anthers ovate, ca. 0.2 mm. |
Fruiting pedicels | divaricate-ascending, straight or slightly curved upward, 4–11(–16) mm, pubescent, trichomes simple and 2–4-rayed. |
divaricate-ascending, straight, 1–4(–8) mm, glabrous. |
Fruits | oblong or ovate, plane, flattened, 6–12 × 3.5–5.5 mm; valves pubescent or puberulent, trichomes simple, 0.1–0.4 mm, (sometimes with short-stalked, 2- or 3-rayed ones); ovules 12–24 per ovary; style 0.6–1 mm (stigma distinctly wider than style). |
ovate to oblong, plane, flattened, 4–7.5 × 1.7–3 mm; valves glabrous; ovules 12–16 per ovary; style 0.1–0.3(–0.5) mm. |
Seeds | (brown), ovoid, 1–1.3 × 0.6–0.9 mm. |
ovoid, 0.7–1 × 0.5–0.6 mm. |
2n | = 128, 144. |
= 32. |
Draba corymbosa |
Draba porsildii |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Jul. |
Habitat | Moist tundra, among calcareous or dolomitic rocks, gravel beaches, silt and clay terraces | Rock outcrops, talus, meadows, gravel slopes |
Elevation | 0-1700 m (0-5600 ft) | 600-3000 m (2000-9800 ft) |
Distribution |
AK; BC; NT; NU; YT; Greenland; Europe (Norway, n Russia); e Asia (Russian Far East, Siberia) |
AK; MT; WY; AB; YT |
Discussion | Draba corymbosa, which is 16-ploid or 18-ploid with x = 8, is an extremely variable species of polyphyletic, allopolyploid origin. Most individuals appear to have decaploid D. alpina (2n = 80) in their parentage, but the other genomes are provided by hexaploid (2n = 48) and octoploid (2n = 64) species (C. Brochmann et al. 1993). O. E. Schulz reduced D. corymbosa to a variety of D. alpina; R. C. Rollins (1993) treated it as a distinct species. Draba corymbosa is distinguished from D. alpina by having pubescent or puberulent (versus glabrous or glabrescent) fruits, corymbose (versus usually elongated) fruiting racemes, and abaxial leaf blade surfaces always lacking (versus usually with some) simple trichomes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
According to H. H. Grundt et al. (2004), tetraploid Draba porsildii probably originated from a diploid ancestor such as D. lonchocarpa. The Wyoming plants are a close match to the type of the species, though they have distinctly shorter pedicels. Some of the Alaskan plants are more problematic; they have finer and more branched trichomes, larger flowers, longer pedicels, and longer styles than the type. It is likely that they belong to another species; detailed molecular and cytological studies on this complex are needed before any meaningful conclusions are reached. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 301. | FNA vol. 7, p. 331. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. alpina var. bellii, D. alpina var. corymbosa, D. barbata, D. bellii, D. kjellmanii, D. macrocarpa, D. vestita | D. nivalis var. brevicula, D. porsildii var. brevicula |
Name authority | R. Brown ex de Candolle: Syst. Nat. 2: 343. (1821) | G. A. Mulligan: Canad. J. Bot. 52: 1795, fig. 8. (1974) |
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