Draba corymbosa |
Draba palanderiana |
|
---|---|---|
Baffin bay Draba, Baffin bay whitlow-grass, flat-top Draba |
palander's Draba, palander's whitlow-grass |
|
Habit | Perennials; (cespitose); caudex branched (with persistent leaves or leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (cespitose); caudex branched (with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, (0.05–)0.2–0.8(–1.5) dm, pubescent throughout, trichomes simple and 2-rayed, 0.4–1 mm, with 3–5-rayed ones, 0.05–0.4 mm, (sometimes trichomes mostly simple). |
unbranched, 0.2–1.1(–1.5) dm, pubescent proximally, trichomes 2–9-rayed, 0.05–0.2 mm, glabrous or sparsely pubescent distally. |
Basal leaves | rosulate; petiolate; petiole base and margin ciliate, (trichomes often course, simple, 0.4–1.3 mm); blade oblanceolate to obovate, 0.6–1.8 cm × 1.5–5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–6-rayed trichomes, 0.2–0.6 mm, adaxially with primarily simple and stalked, 2-rayed trichomes, to 1.1 mm, with 3–5-rayed ones, 0.2–0.4 mm. |
rosulate; petiolate; petiole (thickened), ciliate proximally, (margin not ciliate, trichomes simple and 2-rayed, 0.2–0.5 mm); blade oblanceolate to obovate, 0.3–0.9(–1.5) cm × 1–4 mm, margins usually entire, rarely denticulate, surfaces pubescent with minutely stalked, 8–16-rayed, stellate trichomes, 0.08–0.3 mm, (midvein obscure abaxially). |
Cauline leaves | 0. |
0. |
Racemes | 2–9(–12)-flowered, ebracteate, (corymbose), slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
5–17-flowered, ebracteate, elongated in fruit; rachis not flexuous, usually glabrous, rarely sparsely pubescent as stem. |
Flowers | sepals (grayish green), broadly ovate, 2.2–3 mm, pubescent, (trichomes simple, to 1 mm, sometimes with stalked, smaller, 2–4-rayed ones); petals (broadly patent), yellow, obovate, 4–6 × 3–5 mm; anthers ovate, 0.3–0.4 mm. |
sepals ovate, 2–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–6-rayed); petals white or creamy white, obovate, 4.5–5.5 × 2–3 mm; anthers ovate, 0.4–0.6 mm. |
Fruiting pedicels | divaricate-ascending, straight or slightly curved upward, 4–11(–16) mm, pubescent, trichomes simple and 2–4-rayed. |
divaricate-ascending, straight, (slender), 4–10 mm, usually glabrous, rarely sparsely pubescent as stem. |
Fruits | oblong or ovate, plane, flattened, 6–12 × 3.5–5.5 mm; valves pubescent or puberulent, trichomes simple, 0.1–0.4 mm, (sometimes with short-stalked, 2- or 3-rayed ones); ovules 12–24 per ovary; style 0.6–1 mm (stigma distinctly wider than style). |
(often aborting, becoming lopsided), oblong to elliptic or ovate, plane, flattened, 4–8 × 2–3 mm; valves glabrous; ovules 18–32 per ovary; style 0.3 0.8 mm. |
Seeds | (brown), ovoid, 1–1.3 × 0.6–0.9 mm. |
ovoid, 0.9–1.1 × 0.5–0.7 mm. |
2n | = 128, 144. |
= 16, 32, 64. |
Draba corymbosa |
Draba palanderiana |
|
Phenology | Flowering Jun–Aug. | Flowering Jun–Jul. |
Habitat | Moist tundra, among calcareous or dolomitic rocks, gravel beaches, silt and clay terraces | Rock outcrops, talus, Dryas fellfields, tundra |
Elevation | 0-1700 m (0-5600 ft) | 0-1800 m (0-5900 ft) |
Distribution |
AK; BC; NT; NU; YT; Greenland; Europe (Norway, n Russia); e Asia (Russian Far East, Siberia) |
AK; NT; YT; e Asia (Russian Far East, Siberia) |
Discussion | Draba corymbosa, which is 16-ploid or 18-ploid with x = 8, is an extremely variable species of polyphyletic, allopolyploid origin. Most individuals appear to have decaploid D. alpina (2n = 80) in their parentage, but the other genomes are provided by hexaploid (2n = 48) and octoploid (2n = 64) species (C. Brochmann et al. 1993). O. E. Schulz reduced D. corymbosa to a variety of D. alpina; R. C. Rollins (1993) treated it as a distinct species. Draba corymbosa is distinguished from D. alpina by having pubescent or puberulent (versus glabrous or glabrescent) fruits, corymbose (versus usually elongated) fruiting racemes, and abaxial leaf blade surfaces always lacking (versus usually with some) simple trichomes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
North American plants of Draba palanderiana are diploid and appear to be self-incompatible, an unusual situation among arctic members of the genus. By contrast, some plants from the Russian Far East are tetraploid (2n = 32) or octoploid (2n = 64) and appear to be self-compatible. Detailed studies are needed to establish whether one or two taxa are involved. Many North American collections of Draba palanderiana have been misidentified as D. nivalis. The species is easily separated from D. nivalis by having ciliate (versus non-ciliate) bases of basal leaves, larger petals (4.5–5.5 × 2–3 versus 2–3.5 × 0.8–1.4 mm) that are pale yellow or cream (versus white), and longer fruiting pedicels 4–10 [versus 1–4.5(–8)] mm. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 301. | FNA vol. 7, p. 326. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. alpina var. bellii, D. alpina var. corymbosa, D. barbata, D. bellii, D. kjellmanii, D. macrocarpa, D. vestita | |
Name authority | R. Brown ex de Candolle: Syst. Nat. 2: 343. (1821) | Kjellman: in A. E. Nordenskiöld, Vega Exp. Vetensk. Iakttag. 2: 45. (1883) |
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