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Baffin bay Draba, Baffin bay whitlow-grass, flat-top Draba

Norwegian Draba

Habit Perennials; (cespitose); caudex branched (with persistent leaves or leaf remains, branches sometimes terminating in sterile rosettes); scapose. Perennials; (cespitose); caudex branched (sometimes with persistent leaf remains); often scapose.
Stems

unbranched, (0.05–)0.2–0.8(–1.5) dm, pubescent throughout, trichomes simple and 2-rayed, 0.4–1 mm, with 3–5-rayed ones, 0.05–0.4 mm, (sometimes trichomes mostly simple).

branched or unbranched, (0.2–)0.4–1.4(–2) dm, pubescent throughout, trichomes simple and 2–4-rayed, 0.2–0.5 mm.

Basal leaves

rosulate; petiolate;

petiole base and margin ciliate, (trichomes often course, simple, 0.4–1.3 mm);

blade oblanceolate to obovate, 0.6–1.8 cm × 1.5–5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–6-rayed trichomes, 0.2–0.6 mm, adaxially with primarily simple and stalked, 2-rayed trichomes, to 1.1 mm, with 3–5-rayed ones, 0.2–0.4 mm.

rosulate; petiolate;

petiole (0–0.5 cm), ciliate throughout;

blade oblanceolate to narrowly obovate or narrowly lanceolate, 0.4–2.1 cm × 1.5–7 mm, margins with 1–3 teeth on each side, (ciliate, trichomes simple and 2-rayed, 0.2–0.8 mm), surfaces pubescent abaxially with stalked, (2–)4(–6)-rayed trichomes, 0.1–0.4 mm, adaxially glabrous or pubescent, with simple and stalked, 2-rayed trichomes, 0.1–0.5 mm.

Cauline leaves

0.

0–3(–5);

sessile;

blade ovate to oblong, margins often dentate, (ciliate).

Racemes

2–9(–12)-flowered, ebracteate, (corymbose), slightly elongated in fruit;

rachis not flexuous, pubescent as stem.

5–23-flowered, ebracteate or proximalmost 1(–3) flowers bracteate, considerably elongated in fruit;

rachis slightly or not flexuous, pubescent as stem.

Flowers

sepals (grayish green), broadly ovate, 2.2–3 mm, pubescent, (trichomes simple, to 1 mm, sometimes with stalked, smaller, 2–4-rayed ones);

petals (broadly patent), yellow, obovate, 4–6 × 3–5 mm;

anthers ovate, 0.3–0.4 mm.

sepals ovate, 1.7–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed);

petals white, spatulate, 2.5–4 × 1.7–2.5 mm;

anthers ovate, 0.3–0.4 mm.

Fruiting pedicels

divaricate-ascending, straight or slightly curved upward, 4–11(–16) mm, pubescent, trichomes simple and 2–4-rayed.

suberect to ascending (often appressed to rachis), straight, (1.5–)2.5–5(–9) mm, pubescent, trichomes simple and 2–4-rayed.

Fruits

oblong or ovate, plane, flattened, 6–12 × 3.5–5.5 mm;

valves pubescent or puberulent, trichomes simple, 0.1–0.4 mm, (sometimes with short-stalked, 2- or 3-rayed ones);

ovules 12–24 per ovary;

style 0.6–1 mm (stigma distinctly wider than style).

(sometimes subappressed to rachis), oblong to lanceolate-elliptic, plane, flattened, 4–8 × 2–3 mm;

valves glabrous or pubescent, trichomes simple and short-stalked, 2-rayed, 0.05–0.25 mm;

ovules 18–26 per ovary;

style (0.01–)0.1–0.4(–0.5) mm, (glabrous, stigma about as wide as style).

Seeds

(brown), ovoid, 1–1.3 × 0.6–0.9 mm.

oblong, 0.9–1.2 × 0.5–0.7 mm.

2n

= 128, 144.

= 48.

Draba corymbosa

Draba norvegica

Phenology Flowering Jun–Aug. Flowering Jun–Aug.
Habitat Moist tundra, among calcareous or dolomitic rocks, gravel beaches, silt and clay terraces Rock outcrops and sea cliffs, talus, gravelly and sandy terraces, moist bluffs, turfy limestone shores
Elevation 0-1700 m (0-5600 ft) 0-500 m (0-1600 ft)
Distribution
from FNA
AK; BC; NT; NU; YT; Greenland; Europe (Norway, n Russia); e Asia (Russian Far East, Siberia)
[BONAP county map]
from FNA
NL; NS; NT; QC; Greenland; Europe (Finland, Norway, n Russia, Scotland, Sweden); Atlantic Islands (Iceland)
[BONAP county map]
Discussion

Draba corymbosa, which is 16-ploid or 18-ploid with x = 8, is an extremely variable species of polyphyletic, allopolyploid origin. Most individuals appear to have decaploid D. alpina (2n = 80) in their parentage, but the other genomes are provided by hexaploid (2n = 48) and octoploid (2n = 64) species (C. Brochmann et al. 1993). O. E. Schulz reduced D. corymbosa to a variety of D. alpina; R. C. Rollins (1993) treated it as a distinct species. Draba corymbosa is distinguished from D. alpina by having pubescent or puberulent (versus glabrous or glabrescent) fruits, corymbose (versus usually elongated) fruiting racemes, and abaxial leaf blade surfaces always lacking (versus usually with some) simple trichomes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba norvegica is a highly variable hexaploid taxon and, as delimited herein, it probably represents two taxa with the same distribution and same chromosome number. It is related to hexaploid D. arctogena, from which it is distinguished by characteristics listed under 8. D. arctogena.

Draba norvegica was divided by O. E. Schulz (1927) into eight varieties, whereas R. C. Rollins (1993) recognized just two. In the latter treatment, var. clivicola is distinguished by having predominantly branched, appressed trichomes on stems proximally, lanceolate, glabrous fruits, and slender, erect-appressed pedicels. By contrast, var. norvegica is said to have predominantly simple, spreading trichomes on stems proximally, narrowly ovate to elliptical, glabrous or pubescent fruits, and stout, divaricately ascending pedicels. These differences are extremes of a continuum, and we agree with G. A. Mulligan (1970) in reducing D. clivicola to synonymy of D. norvegica. Rollins indicated that D. norvegica grows in Minnesota, but we have not seen any material from that state.

Draba norvegica is found in Europe in arctic and subarctic areas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 301. FNA vol. 7, p. 322.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms D. alpina var. bellii, D. alpina var. corymbosa, D. barbata, D. bellii, D. kjellmanii, D. macrocarpa, D. vestita D. clivicola, D. hirta var. norvegica, D. norvegica var. clivicola, D. norvegica var. hebecarpa, D. rupestris, D. scandinavica var. hebecarpa, D. trichella
Name authority R. Brown ex de Candolle: Syst. Nat. 2: 343. (1821) Gunnerus: Fl. Norveg. 2: 106. (1772)
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