Draba corymbosa |
Draba mogollonica |
|
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Baffin bay Draba, Baffin bay whitlow-grass, flat-top Draba |
Mogollon Mountain Draba |
|
Habit | Perennials; (cespitose); caudex branched (with persistent leaves or leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Annuals, biennials, or, rarely, perennials; (short-lived); not scapose. |
Stems | unbranched, (0.05–)0.2–0.8(–1.5) dm, pubescent throughout, trichomes simple and 2-rayed, 0.4–1 mm, with 3–5-rayed ones, 0.05–0.4 mm, (sometimes trichomes mostly simple). |
usually branched, (0.6–)1.2–2.8(–3.6) dm, pubescent proximally, glabrous distally, trichomes simple, 0.3–1.3 mm, and 2–4-rayed, 0.1–0.6 mm. |
Basal leaves | rosulate; petiolate; petiole base and margin ciliate, (trichomes often course, simple, 0.4–1.3 mm); blade oblanceolate to obovate, 0.6–1.8 cm × 1.5–5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–6-rayed trichomes, 0.2–0.6 mm, adaxially with primarily simple and stalked, 2-rayed trichomes, to 1.1 mm, with 3–5-rayed ones, 0.2–0.4 mm. |
rosulate; petiole (0–2 cm), densely pubescent, (trichomes simple and 2–4-rayed, not ciliate); blade spatulate to obovate or oblanceolate, (1.5–)3–8(–10) cm × 7–25(–33) mm, margins dentate to denticulate, surfaces pubescent with long-stalked, cruciform, and fewer 2- or 3-rayed trichomes, 0.1–0.6 mm. |
Cauline leaves | 0. |
1–3; sessile; blade oblong to ovate, margins entire, surfaces pubescent as basal. |
Racemes | 2–9(–12)-flowered, ebracteate, (corymbose), slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
12–40(–51)-flowered, ebracteate, elongated in fruit; rachis not flexuous, usually glabrous, rarely proximally with relatively few trichomes. |
Flowers | sepals (grayish green), broadly ovate, 2.2–3 mm, pubescent, (trichomes simple, to 1 mm, sometimes with stalked, smaller, 2–4-rayed ones); petals (broadly patent), yellow, obovate, 4–6 × 3–5 mm; anthers ovate, 0.3–0.4 mm. |
sepals oblong, 2.5–3.5 mm, sparsely pubescent, (trichomes simple and 2–4-rayed); petals yellow, spatulate, (5–)6–8.5 × 2.2–3.5 mm; anthers oblong, 0.7–0.9 mm. |
Fruiting pedicels | divaricate-ascending, straight or slightly curved upward, 4–11(–16) mm, pubescent, trichomes simple and 2–4-rayed. |
horizontal to divaricate, straight, (6–)9–18(–22) mm, usually glabrous, rarely sparsely pubescent as stem proximally. |
Fruits | oblong or ovate, plane, flattened, 6–12 × 3.5–5.5 mm; valves pubescent or puberulent, trichomes simple, 0.1–0.4 mm, (sometimes with short-stalked, 2- or 3-rayed ones); ovules 12–24 per ovary; style 0.6–1 mm (stigma distinctly wider than style). |
linear-elliptic to elliptic-lanceolate, slightly twisted, flattened, 6–19 × 2.5–3.5 mm; valves glabrous or pubescent, trichomes simple and short-stalked, 2(–4)-rayed, 0.07–0.3 mm; ovules 24–36 per ovary; style (0.8–)1–2.2 mm. |
Seeds | (brown), ovoid, 1–1.3 × 0.6–0.9 mm. |
oblong, 1–1.4 × 0.7–0.8 mm. |
2n | = 128, 144. |
= 22. |
Draba corymbosa |
Draba mogollonica |
|
Phenology | Flowering Jun–Aug. | Flowering Apr–May. |
Habitat | Moist tundra, among calcareous or dolomitic rocks, gravel beaches, silt and clay terraces | Rock outcrops and rocky slopes in oak-pine woodlands and mixed conifer communities |
Elevation | 0-1700 m (0-5600 ft) | 1500-2300 m (4900-7500 ft) |
Distribution |
AK; BC; NT; NU; YT; Greenland; Europe (Norway, n Russia); e Asia (Russian Far East, Siberia) |
AZ; NM |
Discussion | Draba corymbosa, which is 16-ploid or 18-ploid with x = 8, is an extremely variable species of polyphyletic, allopolyploid origin. Most individuals appear to have decaploid D. alpina (2n = 80) in their parentage, but the other genomes are provided by hexaploid (2n = 48) and octoploid (2n = 64) species (C. Brochmann et al. 1993). O. E. Schulz reduced D. corymbosa to a variety of D. alpina; R. C. Rollins (1993) treated it as a distinct species. Draba corymbosa is distinguished from D. alpina by having pubescent or puberulent (versus glabrous or glabrescent) fruits, corymbose (versus usually elongated) fruiting racemes, and abaxial leaf blade surfaces always lacking (versus usually with some) simple trichomes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba mogollonica is distributed in New Mexico primarily in the Mogollon and neighboring mountains in Catron, Grant, Sierra, and Socorro counties. It is reported herein for the first time from Arizona from Greenlee County, based on Barneby 2297 (NY), Rollins & Rollins 93113 (CAS, GH), and Windham 2918 (MO, UT). The only published chromosome count (n = 16, compiled in S. I. Warwick and I. A. Al-Shehbaz 2006) is erroneous; counts by M. D. Windham (unpubl.) from five populations representing all of the above-mentioned counties in Arizona and New Mexico consistently agree on n = 11. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 301. | FNA vol. 7, p. 320. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. alpina var. bellii, D. alpina var. corymbosa, D. barbata, D. bellii, D. kjellmanii, D. macrocarpa, D. vestita | |
Name authority | R. Brown ex de Candolle: Syst. Nat. 2: 343. (1821) | Greene: Bot. Gaz. 6: 157. (1881) |
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