Draba corymbosa |
Draba crassa |
|
---|---|---|
Baffin bay Draba, Baffin bay whitlow-grass, flat-top Draba |
thickleaf Draba |
|
Habit | Perennials; (cespitose); caudex branched (with persistent leaves or leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (cespitose); caudex simple or branched (well-developed, with persistent leaf bases); not scapose. |
Stems | unbranched, (0.05–)0.2–0.8(–1.5) dm, pubescent throughout, trichomes simple and 2-rayed, 0.4–1 mm, with 3–5-rayed ones, 0.05–0.4 mm, (sometimes trichomes mostly simple). |
(decumbent to ascending), unbranched, (0.4–)0.6–1.3(–1.5) dm, usually glabrous proximally, rarely sparsely pubescent, pubescent distally, trichomes simple and short-stalked, 2-rayed, (crisped), 0.1–0.6(–0.8) mm. |
Basal leaves | rosulate; petiolate; petiole base and margin ciliate, (trichomes often course, simple, 0.4–1.3 mm); blade oblanceolate to obovate, 0.6–1.8 cm × 1.5–5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–6-rayed trichomes, 0.2–0.6 mm, adaxially with primarily simple and stalked, 2-rayed trichomes, to 1.1 mm, with 3–5-rayed ones, 0.2–0.4 mm. |
rosulate; petiolate; petiole ciliate, (trichomes mostly simple, 0.3–0.8 mm); blade oblanceolate, 2–6(–7) cm × 2.5–8(–10) mm, margins entire, (pubescent as petiole), surfaces glabrous. |
Cauline leaves | 0. |
2–4(–6); sessile; blade ovate to oblong, margins entire, (ciliate, trichomes simple and stalked, 2-rayed). |
Racemes | 2–9(–12)-flowered, ebracteate, (corymbose), slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
(4–)8–20(–25)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent, trichomes simple and short-stalked, 2-rayed, (crisped), (0.1–0.8 mm). |
Flowers | sepals (grayish green), broadly ovate, 2.2–3 mm, pubescent, (trichomes simple, to 1 mm, sometimes with stalked, smaller, 2–4-rayed ones); petals (broadly patent), yellow, obovate, 4–6 × 3–5 mm; anthers ovate, 0.3–0.4 mm. |
sepals (green tinged purplish), ovate to oblong, 2–3(–3.3) mm, pubescent, (trichomes simple and 2-rayed); petals yellow, spatulate to subobovate, 3.5–5(–6) × (2–) 2.5–3.5(–4) mm; anthers ovate, 0.5–0.6 mm. |
Fruiting pedicels | divaricate-ascending, straight or slightly curved upward, 4–11(–16) mm, pubescent, trichomes simple and 2–4-rayed. |
divaricate-ascending, straight, 5–10(–15) mm, pubescent as rachis. |
Fruits | oblong or ovate, plane, flattened, 6–12 × 3.5–5.5 mm; valves pubescent or puberulent, trichomes simple, 0.1–0.4 mm, (sometimes with short-stalked, 2- or 3-rayed ones); ovules 12–24 per ovary; style 0.6–1 mm (stigma distinctly wider than style). |
lanceolate to ovate-lanceolate, slightly twisted, flattened, (7–)8–14 × 3–5 mm; valves glabrous; ovules 16–20 per ovary; style (0.4–)0.7–1.2(–1.5) mm. |
Seeds | (brown), ovoid, 1–1.3 × 0.6–0.9 mm. |
oblong, 1.2–1.7 × 0.8–1.1 mm. |
2n | = 128, 144. |
= 24. |
Draba corymbosa |
Draba crassa |
|
Phenology | Flowering Jun–Aug. | Flowering Jul–Aug. |
Habitat | Moist tundra, among calcareous or dolomitic rocks, gravel beaches, silt and clay terraces | Rock outcrops and talus, alpine tundra, rocky meadows |
Elevation | 0-1700 m (0-5600 ft) | 2900-4300 m (9500-14100 ft) |
Distribution |
AK; BC; NT; NU; YT; Greenland; Europe (Norway, n Russia); e Asia (Russian Far East, Siberia) |
CO; MT; NM; UT; WY
|
Discussion | Draba corymbosa, which is 16-ploid or 18-ploid with x = 8, is an extremely variable species of polyphyletic, allopolyploid origin. Most individuals appear to have decaploid D. alpina (2n = 80) in their parentage, but the other genomes are provided by hexaploid (2n = 48) and octoploid (2n = 64) species (C. Brochmann et al. 1993). O. E. Schulz reduced D. corymbosa to a variety of D. alpina; R. C. Rollins (1993) treated it as a distinct species. Draba corymbosa is distinguished from D. alpina by having pubescent or puberulent (versus glabrous or glabrescent) fruits, corymbose (versus usually elongated) fruiting racemes, and abaxial leaf blade surfaces always lacking (versus usually with some) simple trichomes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba crassa is easily recognized by its combination of fleshy, distinctly petiolate leaves with glabrous surfaces and mostly decumbent flowering/fruiting stems. N. H. Holmgren (2005b) indicated that it has 52 seeds per fruit (26 per locule), but in the plants we examined, the seed count did not exceed 20 per fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 301. | FNA vol. 7, p. 302. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. alpina var. bellii, D. alpina var. corymbosa, D. barbata, D. bellii, D. kjellmanii, D. macrocarpa, D. vestita | D. chrysantha var. crassa |
Name authority | R. Brown ex de Candolle: Syst. Nat. 2: 343. (1821) | Rydberg: Mem. New York Bot. Gard. 1: 182. (1900) |
Web links |