Draba corymbosa |
Draba bifurcata |
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Baffin bay Draba, Baffin bay whitlow-grass, flat-top Draba |
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Habit | Perennials; (cespitose); caudex branched (with persistent leaves or leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Biennials or, rarely, perennials; caudex simple or branched; not scapose. |
Stems | unbranched, (0.05–)0.2–0.8(–1.5) dm, pubescent throughout, trichomes simple and 2-rayed, 0.4–1 mm, with 3–5-rayed ones, 0.05–0.4 mm, (sometimes trichomes mostly simple). |
branched or unbranched, 1–3.9 dm, pubescent throughout or proximally, trichomes simple, 0.3–1.4 mm, often with smaller, 2 (or 3)-rayed ones. |
Basal leaves | rosulate; petiolate; petiole base and margin ciliate, (trichomes often course, simple, 0.4–1.3 mm); blade oblanceolate to obovate, 0.6–1.8 cm × 1.5–5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–6-rayed trichomes, 0.2–0.6 mm, adaxially with primarily simple and stalked, 2-rayed trichomes, to 1.1 mm, with 3–5-rayed ones, 0.2–0.4 mm. |
(soon withered); rosulate; petiolate; petiole ciliate, (trichomes simple, 0.4–1.1 mm); blade oblanceolate to spatulate, 1–7 cm × 2–12(–26) mm, margins entire or dentate (pubescent as petiole), surfaces pubescent, abaxially with simple trichomes, 0.6–1.1 mm, and stalked, 2- (or 3-)rayed ones, often smaller, adaxially with subsetiform, simple trichomes, 0.5–1.2 mm (rarely with fewer, stalked, 2-rayed ones). |
Cauline leaves | 0. |
(2–)6–15; sessile; blade lanceolate to ovate or oblong, margins dentate, surfaces pubescent as basal. |
Racemes | 2–9(–12)-flowered, ebracteate, (corymbose), slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
10–51-flowered, ebracteate, elongated in fruit; rachis glabrous or pubescent as stem, not flexuous. |
Flowers | sepals (grayish green), broadly ovate, 2.2–3 mm, pubescent, (trichomes simple, to 1 mm, sometimes with stalked, smaller, 2–4-rayed ones); petals (broadly patent), yellow, obovate, 4–6 × 3–5 mm; anthers ovate, 0.3–0.4 mm. |
sepals ovate, 2.5–3.2 mm, pubescent, (trichomes simple and short-stalked, 2-rayed); petals yellow, oblanceolate, 4.5–6.5 × 1.5–2 mm; anthers oblong, 0.8–1.1 mm. |
Fruiting pedicels | divaricate-ascending, straight or slightly curved upward, 4–11(–16) mm, pubescent, trichomes simple and 2–4-rayed. |
horizontal to divaricate, straight, 4–15 mm, glabrous or pubescent, trichomes simple and 2-rayed. |
Fruits | oblong or ovate, plane, flattened, 6–12 × 3.5–5.5 mm; valves pubescent or puberulent, trichomes simple, 0.1–0.4 mm, (sometimes with short-stalked, 2- or 3-rayed ones); ovules 12–24 per ovary; style 0.6–1 mm (stigma distinctly wider than style). |
elliptic-lanceolate, twisted (1 turn or plane), flattened, 6–10 × 2–2.5 mm; valves glabrous or sparsely puberulent, trichomes simple, 0.05–0.15 mm; ovules 14–20 per ovary; style (1–)2–3 mm. |
Seeds | (brown), ovoid, 1–1.3 × 0.6–0.9 mm. |
oblong, 1.1–1.4 × 0.7–0.9 mm. |
2n | = 128, 144. |
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Draba corymbosa |
Draba bifurcata |
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Phenology | Flowering Jun–Aug. | Flowering Jun–Jul. |
Habitat | Moist tundra, among calcareous or dolomitic rocks, gravel beaches, silt and clay terraces | Rocky areas, damp shady ravines, aspen-spruce communities |
Elevation | 0-1700 m (0-5600 ft) | 1800-3600 m (5900-11800 ft) |
Distribution |
AK; BC; NT; NU; YT; Greenland; Europe (Norway, n Russia); e Asia (Russian Far East, Siberia) |
AZ |
Discussion | Draba corymbosa, which is 16-ploid or 18-ploid with x = 8, is an extremely variable species of polyphyletic, allopolyploid origin. Most individuals appear to have decaploid D. alpina (2n = 80) in their parentage, but the other genomes are provided by hexaploid (2n = 48) and octoploid (2n = 64) species (C. Brochmann et al. 1993). O. E. Schulz reduced D. corymbosa to a variety of D. alpina; R. C. Rollins (1993) treated it as a distinct species. Draba corymbosa is distinguished from D. alpina by having pubescent or puberulent (versus glabrous or glabrescent) fruits, corymbose (versus usually elongated) fruiting racemes, and abaxial leaf blade surfaces always lacking (versus usually with some) simple trichomes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba bifurcata is distinguished from other North American species by a combination of yellow flowers with styles (1–)2–3 mm and almost exclusively simple trichomes on the abaxial surface of cauline leaf blades. It is known only from Chiricahua, Santa Catalina, and White mountains in Apache, Cochise, and Pima counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 301. | FNA vol. 7, p. 295. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. alpina var. bellii, D. alpina var. corymbosa, D. barbata, D. bellii, D. kjellmanii, D. macrocarpa, D. vestita | D. helleriana var. bifurcata |
Name authority | R. Brown ex de Candolle: Syst. Nat. 2: 343. (1821) | (C. L. Hitchcock) Al-Shehbaz & Windham: Harvard Pap. Bot. 12: 411. (2007) |
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