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Baffin bay Draba, Baffin bay whitlow-grass, flat-top Draba

rough Draba, rough whitlowgrass

Habit Perennials; (cespitose); caudex branched (with persistent leaves or leaf remains, branches sometimes terminating in sterile rosettes); scapose. Perennials; (cespitose); caudex branched (with some persistent leaf bases); scapose.
Stems

unbranched, (0.05–)0.2–0.8(–1.5) dm, pubescent throughout, trichomes simple and 2-rayed, 0.4–1 mm, with 3–5-rayed ones, 0.05–0.4 mm, (sometimes trichomes mostly simple).

unbranched, (0.4–)0.6–1.5(–2.7) dm, pubescent throughout, trichomes simple, 0.5–1.5 mm, and 2–4-rayed ones, 0.05–0.3 mm, (sometimes 2–4-rayed ones distally).

Basal leaves

rosulate; petiolate;

petiole base and margin ciliate, (trichomes often course, simple, 0.4–1.3 mm);

blade oblanceolate to obovate, 0.6–1.8 cm × 1.5–5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–6-rayed trichomes, 0.2–0.6 mm, adaxially with primarily simple and stalked, 2-rayed trichomes, to 1.1 mm, with 3–5-rayed ones, 0.2–0.4 mm.

rosulate;

subsessile or shortly petiolate;

petiole ciliate throughout;

blade oblanceolate to spatulate or obovate, (0.6–)0.8–4.5(–6) cm × 2–12(–16) mm, margins usually entire, rarely obscurely dentate, (not ciliate), surfaces abaxially pubescent with stalked, (2–)4-rayed trichomes, 0.2–1 mm, adaxially with stalked, 2–4-rayed trichomes, 0.3–0.8 mm, and simple ones, 0.5–1.9 mm.

Cauline leaves

0.

0.

Racemes

2–9(–12)-flowered, ebracteate, (corymbose), slightly elongated in fruit;

rachis not flexuous, pubescent as stem.

(15–)30–75-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent, trichomes 2–4-rayed, sometimes with simple ones.

Flowers

sepals (grayish green), broadly ovate, 2.2–3 mm, pubescent, (trichomes simple, to 1 mm, sometimes with stalked, smaller, 2–4-rayed ones);

petals (broadly patent), yellow, obovate, 4–6 × 3–5 mm;

anthers ovate, 0.3–0.4 mm.

sepals broadly ovate, 1.8–3.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed);

petals pale yellow, oblanceolate, 4–7 × 1.5–2.5 mm;

anthers ovate to oblong, 0.4–0.7 mm.

Fruiting pedicels

divaricate-ascending, straight or slightly curved upward, 4–11(–16) mm, pubescent, trichomes simple and 2–4-rayed.

horizontal to divaricate-ascending, straight, 3–11(–16) mm, pubescent, trichomes 2–4-rayed, sometimes with simple ones.

Fruits

oblong or ovate, plane, flattened, 6–12 × 3.5–5.5 mm;

valves pubescent or puberulent, trichomes simple, 0.1–0.4 mm, (sometimes with short-stalked, 2- or 3-rayed ones);

ovules 12–24 per ovary;

style 0.6–1 mm (stigma distinctly wider than style).

ovoid-ellipsoid to oblong, plane, inflated basally, 3–8 × 1.8–3.5 mm;

valves hirsute, trichomes simple and spurred, (0.2–)0.3–1 mm, or puberulent, trichomes short-stalked, 2–4-rayed, 0.1–0.2 mm;

ovules 8–12(–18) per ovary;

style 0.6–2(–2.5) mm.

Seeds

(brown), ovoid, 1–1.3 × 0.6–0.9 mm.

oblong, 0.9–1.2 × 0.6–0.8 mm.

2n

= 128, 144.

= 30.

Draba corymbosa

Draba asprella

Phenology Flowering Jun–Aug.
Habitat Moist tundra, among calcareous or dolomitic rocks, gravel beaches, silt and clay terraces
Elevation 0-1700 m (0-5600 ft)
Distribution
from FNA
AK; BC; NT; NU; YT; Greenland; Europe (Norway, n Russia); e Asia (Russian Far East, Siberia)
[BONAP county map]
from FNA
AZ
[WildflowerSearch map]
[BONAP county map]
Discussion

Draba corymbosa, which is 16-ploid or 18-ploid with x = 8, is an extremely variable species of polyphyletic, allopolyploid origin. Most individuals appear to have decaploid D. alpina (2n = 80) in their parentage, but the other genomes are provided by hexaploid (2n = 48) and octoploid (2n = 64) species (C. Brochmann et al. 1993). O. E. Schulz reduced D. corymbosa to a variety of D. alpina; R. C. Rollins (1993) treated it as a distinct species. Draba corymbosa is distinguished from D. alpina by having pubescent or puberulent (versus glabrous or glabrescent) fruits, corymbose (versus usually elongated) fruiting racemes, and abaxial leaf blade surfaces always lacking (versus usually with some) simple trichomes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Both R. C. Rollins (1993) and N. H. Holmgren (2005b) indicated that the chromosome number of Draba asprella is 2n = 32, but repeated counts by one of us (M. D. Windham 2000, unpubl.) show that the species consistently has 2n = 30. Rollins divided D. asprella into four varieties encompassing tremendous variation in trichome morphology. One of these (var. zionensis) is more closely related to D. sobolifera (Windham and L. Allphin, unpubl.) and is treated herein at species rank. Within D. asprella, in the strict sense, we recognize two varieties separated primarily by the type of trichomes found on the fruits. The distinctions are not absolute and there appear to be forms connecting the two, especially in Coconino County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruits hirsute, trichomes simple and unequally 2-rayed, (0.2-)0.3-1 mm.
var. asprella
1. Fruits puberulent, trichomes 2-4-rayed, 0.1-0.2 mm.
var. stelligera
Source FNA vol. 7, p. 301. FNA vol. 7, p. 293.
Parent taxa Brassicaceae > tribe Arabideae > Draba Brassicaceae > tribe Arabideae > Draba
Sibling taxa
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Subordinate taxa
D. asprella var. asprella, D. asprella var. stelligera
Synonyms D. alpina var. bellii, D. alpina var. corymbosa, D. barbata, D. bellii, D. kjellmanii, D. macrocarpa, D. vestita
Name authority R. Brown ex de Candolle: Syst. Nat. 2: 343. (1821) Greene: Bull. Torrey Bot. Club 10: 125. (1883)
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