Draba corymbosa |
Draba asprella |
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Baffin bay Draba, Baffin bay whitlow-grass, flat-top Draba |
rough Draba, rough whitlowgrass |
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Habit | Perennials; (cespitose); caudex branched (with persistent leaves or leaf remains, branches sometimes terminating in sterile rosettes); scapose. | Perennials; (cespitose); caudex branched (with some persistent leaf bases); scapose. | ||||
Stems | unbranched, (0.05–)0.2–0.8(–1.5) dm, pubescent throughout, trichomes simple and 2-rayed, 0.4–1 mm, with 3–5-rayed ones, 0.05–0.4 mm, (sometimes trichomes mostly simple). |
unbranched, (0.4–)0.6–1.5(–2.7) dm, pubescent throughout, trichomes simple, 0.5–1.5 mm, and 2–4-rayed ones, 0.05–0.3 mm, (sometimes 2–4-rayed ones distally). |
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Basal leaves | rosulate; petiolate; petiole base and margin ciliate, (trichomes often course, simple, 0.4–1.3 mm); blade oblanceolate to obovate, 0.6–1.8 cm × 1.5–5 mm, margins entire, surfaces pubescent, abaxially with stalked, 2–6-rayed trichomes, 0.2–0.6 mm, adaxially with primarily simple and stalked, 2-rayed trichomes, to 1.1 mm, with 3–5-rayed ones, 0.2–0.4 mm. |
rosulate; subsessile or shortly petiolate; petiole ciliate throughout; blade oblanceolate to spatulate or obovate, (0.6–)0.8–4.5(–6) cm × 2–12(–16) mm, margins usually entire, rarely obscurely dentate, (not ciliate), surfaces abaxially pubescent with stalked, (2–)4-rayed trichomes, 0.2–1 mm, adaxially with stalked, 2–4-rayed trichomes, 0.3–0.8 mm, and simple ones, 0.5–1.9 mm. |
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Cauline leaves | 0. |
0. |
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Racemes | 2–9(–12)-flowered, ebracteate, (corymbose), slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
(15–)30–75-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent, trichomes 2–4-rayed, sometimes with simple ones. |
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Flowers | sepals (grayish green), broadly ovate, 2.2–3 mm, pubescent, (trichomes simple, to 1 mm, sometimes with stalked, smaller, 2–4-rayed ones); petals (broadly patent), yellow, obovate, 4–6 × 3–5 mm; anthers ovate, 0.3–0.4 mm. |
sepals broadly ovate, 1.8–3.5 mm, pubescent, (trichomes simple and short-stalked, 2–4-rayed); petals pale yellow, oblanceolate, 4–7 × 1.5–2.5 mm; anthers ovate to oblong, 0.4–0.7 mm. |
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Fruiting pedicels | divaricate-ascending, straight or slightly curved upward, 4–11(–16) mm, pubescent, trichomes simple and 2–4-rayed. |
horizontal to divaricate-ascending, straight, 3–11(–16) mm, pubescent, trichomes 2–4-rayed, sometimes with simple ones. |
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Fruits | oblong or ovate, plane, flattened, 6–12 × 3.5–5.5 mm; valves pubescent or puberulent, trichomes simple, 0.1–0.4 mm, (sometimes with short-stalked, 2- or 3-rayed ones); ovules 12–24 per ovary; style 0.6–1 mm (stigma distinctly wider than style). |
ovoid-ellipsoid to oblong, plane, inflated basally, 3–8 × 1.8–3.5 mm; valves hirsute, trichomes simple and spurred, (0.2–)0.3–1 mm, or puberulent, trichomes short-stalked, 2–4-rayed, 0.1–0.2 mm; ovules 8–12(–18) per ovary; style 0.6–2(–2.5) mm. |
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Seeds | (brown), ovoid, 1–1.3 × 0.6–0.9 mm. |
oblong, 0.9–1.2 × 0.6–0.8 mm. |
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2n | = 128, 144. |
= 30. |
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Draba corymbosa |
Draba asprella |
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Phenology | Flowering Jun–Aug. | |||||
Habitat | Moist tundra, among calcareous or dolomitic rocks, gravel beaches, silt and clay terraces | |||||
Elevation | 0-1700 m (0-5600 ft) | |||||
Distribution |
AK; BC; NT; NU; YT; Greenland; Europe (Norway, n Russia); e Asia (Russian Far East, Siberia) |
AZ
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Discussion | Draba corymbosa, which is 16-ploid or 18-ploid with x = 8, is an extremely variable species of polyphyletic, allopolyploid origin. Most individuals appear to have decaploid D. alpina (2n = 80) in their parentage, but the other genomes are provided by hexaploid (2n = 48) and octoploid (2n = 64) species (C. Brochmann et al. 1993). O. E. Schulz reduced D. corymbosa to a variety of D. alpina; R. C. Rollins (1993) treated it as a distinct species. Draba corymbosa is distinguished from D. alpina by having pubescent or puberulent (versus glabrous or glabrescent) fruits, corymbose (versus usually elongated) fruiting racemes, and abaxial leaf blade surfaces always lacking (versus usually with some) simple trichomes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Both R. C. Rollins (1993) and N. H. Holmgren (2005b) indicated that the chromosome number of Draba asprella is 2n = 32, but repeated counts by one of us (M. D. Windham 2000, unpubl.) show that the species consistently has 2n = 30. Rollins divided D. asprella into four varieties encompassing tremendous variation in trichome morphology. One of these (var. zionensis) is more closely related to D. sobolifera (Windham and L. Allphin, unpubl.) and is treated herein at species rank. Within D. asprella, in the strict sense, we recognize two varieties separated primarily by the type of trichomes found on the fruits. The distinctions are not absolute and there appear to be forms connecting the two, especially in Coconino County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 301. | FNA vol. 7, p. 293. | ||||
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | D. alpina var. bellii, D. alpina var. corymbosa, D. barbata, D. bellii, D. kjellmanii, D. macrocarpa, D. vestita | |||||
Name authority | R. Brown ex de Candolle: Syst. Nat. 2: 343. (1821) | Greene: Bull. Torrey Bot. Club 10: 125. (1883) | ||||
Web links |