Draba cinerea |
Draba incrassata |
|
---|---|---|
gray-leaf Draba, gray-leaf whitlow-grass |
Sweetwater Draba, Sweetwater Mountains. Draba |
|
Habit | Perennials; (cespitose); caudex simple or branched; rarely scapose. | Perennials; (cespitose, loosely matted); caudex branched (with persistent leaf bases, branches often creeping, sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, (0.3–)0.5–1.6(–2.7) dm, pubescent throughout, trichomes 4–10-rayed, 0.1–0.3 mm, (often some rays branched, simple trichomes sparse, to 0.6 mm). |
unbranched, 0.2–0.8 dm, glabrous throughout. |
Basal leaves | rosulate; petiolate; petiole base and proximalmost margin ciliate, (trichomes simple, 0.2–0.8 mm); blade oblanceolate to narrowly obovate or linear-lanceolate, 0.4–1.5 cm × 1–5 mm, margins usually entire, rarely with 1 tooth on each side, surfaces densely pubescent with minutely stalked, stellate, 8–12-rayed, trichomes 0.1–0.25 mm, (midvein obscure abaxially, apex trichomes simple). |
rosulate; subsessile; petiole ciliate; blade (thick and somewhat fleshy), obovate to spatulate or oblanceolate, 0.3–1 cm × 1.5–4.5 mm, margins entire, (sometimes sparsely ciliate), surfaces glabrous or pubescent proximal to apex, with simple and fewer, short-stalked, 2-rayed trichomes, 0.2–0.7 mm. |
Cauline leaves | 0–3(–5); sessile; blade ovate or oblong to lanceolate, margins entire, surfaces pubescent as basal. |
0. |
Racemes | (3–)5–17(–24)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
8–22-flowered, ebracteate, elongated in fruit; rachis not flexuous, glabrous. |
Flowers | sepals ovate, 1.7–2.5 mm, pubescent, (trichomes simple and short-stalked, 2- or 3-rayed); petals white, spatulate to obovate, 3.5–4.5 × 1.5–2 mm; anthers ovate, 0.3–0.4 mm. |
sepals oblong, 1.7–3 mm, glabrous or pubescent, (trichomes subapical, simple or 2-rayed); petals bright yellow, oblanceolate to spatulate, 3–5 × 1.7–2.5 mm; anthers oblong, 0.4–0.6 mm. |
Fruiting pedicels | divaricate-ascending or ascending, straight, (3–)4–7(–9) mm, pubescent as stem. |
horizontal to divaricate-ascending (distinctly decurrent basally), curved upward, 3–7(–10) mm, glabrous. |
Fruits | oblong to elliptic, plane, slightly flattened, 5–8 × 2–3 mm; valves pubescent, trichomes short-stalked, 2–5-rayed, 0.05–0.3 mm, (some rays branched); ovules 20–36(–42) per ovary; style (0.1–)0.2–1 mm. |
ovate to ovate-lanceolate or narrowly elliptic-lanceolate, plane, flattened, 3–7(–10) × 2.2–4.5 mm; valves usually glabrous, rarely sparsely puberulent, trichomes simple, 0.03–0.1 mm; ovules 8–12 per ovary; style 0.2–0.8 mm. |
Seeds | ovoid, 0.6–0.8 × 0.4–0.6 mm. |
oblong, 1.5–1.8 × 0.8–1.1 mm. |
2n | = 48. |
= 24. |
Draba cinerea |
Draba incrassata |
|
Phenology | Flowering Jun–Jul. | Flowering Jun–Aug. |
Habitat | Rock outcrops, ridges, meadows, gravel beaches, stream banks, alluvial fans | Rocky and gravelly slopes, alpine fellfields |
Elevation | 0-500 m (0-1600 ft) | 2500-3500 m (8200-11500 ft) |
Distribution |
AK; BC; NT; NU; ON; QC; SK; YT; Greenland; Europe (Finland, Norway, n Russia); e Asia (Russian Far East, Siberia) |
CA |
Discussion | Draba cinerea is a polymorphic species in which tetraploid, hexaploid, heptaploid, and octoploid chromosome counts (all based on x = 8) have been reported (e.g., R. C. Rollins 1993). Although we have not seen the type collection, we tentatively consider the taxon to be hexaploid, which is the predominant ploidy level throughout the species range, including North America, and also from its type region in Siberia. Hexaploid Draba cinerea is closely related to decaploid (2n = 80) D. arctica, from which it is distinguished with difficulty. In general, D. cinerea has 1–3(–5)-leaved stems, basal leaf blades usually not terminated with simple trichomes and with obscure midveins abaxially, petals 3.5–4.5 mm, and seeds 0.6–0.8 × 0.4–0.6 mm. By contrast, D. arctica has leafless or 1(–3)-leaved stems, basal leaf blades terminated with simple or 2-rayed trichomes and with midveins distinct abaxially, petals 3.5–6 mm, and seeds (0.8–)0.9–1.1 × (0.6–)0.7–0.8 mm. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although originally treated as a variety of Draba lemmonii, D. incrassata is quite distinct both morphologically (R. A. Price and R. C. Rollins 1991) and chromosomally (M. D. Windham, unpubl.). The species is known from the Sweetwater Mountains in Mono County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 300. | FNA vol. 7, p. 313. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. magellanica subsp. cinerea | D. lemmonii var. incrassata |
Name authority | Adams: Mém Soc. Imp. Naturalistes Moscou 5: 103. (1817) | (Rollins) Rollins & R. A. Price: Harvard Pap. Bot. 1(3): 73. (1991) |
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