FNA: Draba carnosula vs. Draba longisquamosa

	Draba carnosula	Draba longisquamosa
	Mount Eddy Draba, Mt. Eddy Draba	granite Draba
Habit	Perennials; (loosely cespitose); caudex branched (somewhat surculose, with persistent leaf bases, branches sometimes terminating in sterile rosettes); scapose.	Perennials; (cespitose); caudex branched (covered with persistent petiole remains, branches sometimes terminating in sterile rosettes); scapose.
Stems	unbranched, 0.3–1.2 dm, glabrous throughout.	unbranched, 0.2–0.9 dm, hirsute throughout, trichomes simple, 0.2–0.8 mm.
Basal leaves	rosulate; petiolate; petiole (persistent, midvein prominent), ciliate, (trichomes sparse, short-stalked, 2–4-rayed, sometimes with simple ones, 0.2–0.5 mm); blade (somewhat fleshy), oblanceolate or spatulate to obovate, 0.3–1.5 cm × 1.5–5 mm, margins entire, (pubescent as petiole), surfaces glabrous.	rosulate; shortly petiolate; petiole base and margin ciliate, (trichomes simple, 0.1–1 mm); blade oblanceolate to obovate, 0.5–2 cm × 2–5 mm, margins entire, surfaces pubescent with simple trichomes, 0.15–0.8 mm.
Cauline leaves	0.	0.
Racemes	2–6-flowered, ebracteate, elongated in fruit; rachis not flexuous, glabrous.	4–16-flowered, ebracteate, (subumbellate), slightly elongated in fruit; rachis not flexuous, hirsute as stem.
Flowers	sepals ovate, 3–4 mm, glabrous; petals yellow, oblanceolate, 5–7 × 1.3–2 mm; anthers oblong, 0.7–0.9 mm.	sepals (persistent) ovate, 1.5–2.2 mm, pubescent, (trichomes simple); petals (persistent), yellow, spatulate, 3.5–5 × 1.5–2 mm; anthers oblong, 0.5–0.6 mm.
Fruiting pedicels	divaricate-ascending, straight, 3–8 mm, usually glabrous, rarely trichomes simple.	divaricate to ascending (not decurrent basally), straight, 3–7 mm, hirsute as stem.
Fruits	lanceolate to elliptic-lanceolate, plane, strongly flattened, 10–23 × 4–6 mm; valves (each with distinct midvein), glabrous; ovules 8–12 per ovary; style 2–3 mm.	ovate to suborbicular, plane, flattened, 3.5–7 × 2.5–5 mm; valves pubescent, trichomes simple, 0.1–0.3 mm; ovules 10–16 per ovary; style 0.4–1 mm.
Seeds	(broadly winged), orbicular, 3–4.5 in diam.; (wing 1–1.5 mm wide).	ovoid, 1–1.2 × 0.7–0.8 mm.

	Draba carnosula	Draba longisquamosa
Phenology	Flowering Jun–Jul.	Flowering Jul.
Habitat	Open rocky hillsides	Gravelly areas
Elevation	2800-3100 m (9200-10200 ft)	3000-3900 m (9800-12800 ft)
Distribution	CA [BONAP county map]	CA [BONAP county map]
Discussion	Of conservation concern. Although Draba carnosula was reduced by Hitchcock to a variety of D. howellii, it differs significantly by having leafless flowering stems, 2–6-flowered racemes, and broadly winged seeds 3–4.5 mm in diam. Draba howellii almost always has 1–3-leaved stems, (5–)7–18(–25)-flowered racemes, and not winged, distally appendaged seeds 1–1.6 mm in diam. Draba carnosula is known only from a few collections from Mount Eddy in Trinity County and the north side of Mount Shasta in Siskiyou County. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Draba longisquamosa was treated as a synonym of D. lemmonii by both C. L. Hitchcock (1941) and R. C. Rollins (1993). We feel that the two taxa show sufficient morphological divergence and merit recognition as separate species (see I. A. Al-Shehbaz and M. D. Windham 2007 for more detailed discussion). Draba longisquamosa is currently known only from the southern Sierra Nevada in Fresno, Inyo, and Tulare counties. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 7, p. 299.	FNA vol. 7, p. 317.
Parent taxa	Brassicaceae > tribe Arabideae > Draba	Brassicaceae > tribe Arabideae > Draba
Sibling taxa	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms	D. howellii var. carnosula
Name authority	O. E. Schulz: in H. G. A. Engler, Pflanzenr. 89[IV,105]: 82. (1927)	O. E. Schulz: in H. G. A. Engler, Pflanzenr. 89[IV,105]: 94. (1927)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Draba longisquamosa

Mount Eddy Draba, Mt. Eddy Draba

granite Draba

Habit

Perennials; (loosely cespitose); caudex branched (somewhat surculose, with persistent leaf bases, branches sometimes terminating in sterile rosettes); scapose.

Perennials; (cespitose); caudex branched (covered with persistent petiole remains, branches sometimes terminating in sterile rosettes); scapose.

Stems

unbranched, 0.3–1.2 dm, glabrous throughout.

unbranched, 0.2–0.9 dm, hirsute throughout, trichomes simple, 0.2–0.8 mm.

Basal leaves

rosulate;

petiolate;

petiole (persistent, midvein prominent), ciliate, (trichomes sparse, short-stalked, 2–4-rayed, sometimes with simple ones, 0.2–0.5 mm);

blade (somewhat fleshy), oblanceolate or spatulate to obovate, 0.3–1.5 cm × 1.5–5 mm, margins entire, (pubescent as petiole), surfaces glabrous.

rosulate; shortly petiolate;

petiole base and margin ciliate, (trichomes simple, 0.1–1 mm);

blade oblanceolate to obovate, 0.5–2 cm × 2–5 mm, margins entire, surfaces pubescent with simple trichomes, 0.15–0.8 mm.

Cauline leaves

Racemes

2–6-flowered, ebracteate, elongated in fruit;

rachis not flexuous, glabrous.

4–16-flowered, ebracteate, (subumbellate), slightly elongated in fruit;

rachis not flexuous, hirsute as stem.

Flowers

sepals ovate, 3–4 mm, glabrous;

petals yellow, oblanceolate, 5–7 × 1.3–2 mm;

anthers oblong, 0.7–0.9 mm.

sepals (persistent) ovate, 1.5–2.2 mm, pubescent, (trichomes simple);

petals (persistent), yellow, spatulate, 3.5–5 × 1.5–2 mm;

anthers oblong, 0.5–0.6 mm.

Fruiting pedicels

divaricate-ascending, straight, 3–8 mm, usually glabrous, rarely trichomes simple.

divaricate to ascending (not decurrent basally), straight, 3–7 mm, hirsute as stem.

Fruits

lanceolate to elliptic-lanceolate, plane, strongly flattened, 10–23 × 4–6 mm;

valves (each with distinct midvein), glabrous;

ovules 8–12 per ovary;

style 2–3 mm.

ovate to suborbicular, plane, flattened, 3.5–7 × 2.5–5 mm;

valves pubescent, trichomes simple, 0.1–0.3 mm;

ovules 10–16 per ovary;

style 0.4–1 mm.

Seeds

(broadly winged), orbicular, 3–4.5 in diam.; (wing 1–1.5 mm wide).

ovoid, 1–1.2 × 0.7–0.8 mm.

Draba longisquamosa

Phenology

Flowering Jun–Jul.

Flowering Jul.

Habitat

Open rocky hillsides

Gravelly areas

Elevation

2800-3100 m (9200-10200 ft)

3000-3900 m (9800-12800 ft)

Distribution

[BONAP county map]

Discussion

Of conservation concern.

Although Draba carnosula was reduced by Hitchcock to a variety of D. howellii, it differs significantly by having leafless flowering stems, 2–6-flowered racemes, and broadly winged seeds 3–4.5 mm in diam. Draba howellii almost always has 1–3-leaved stems, (5–)7–18(–25)-flowered racemes, and not winged, distally appendaged seeds 1–1.6 mm in diam. Draba carnosula is known only from a few collections from Mount Eddy in Trinity County and the north side of Mount Shasta in Siskiyou County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba longisquamosa was treated as a synonym of D. lemmonii by both C. L. Hitchcock (1941) and R. C. Rollins (1993). We feel that the two taxa show sufficient morphological divergence and merit recognition as separate species (see I. A. Al-Shehbaz and M. D. Windham 2007 for more detailed discussion). Draba longisquamosa is currently known only from the southern Sierra Nevada in Fresno, Inyo, and Tulare counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 7, p. 299.

FNA vol. 7, p. 317.

Parent taxa

Brassicaceae > tribe Arabideae > Draba

Sibling taxa

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

Synonyms

D. howellii var. carnosula

Name authority

O. E. Schulz: in H. G. A. Engler, Pflanzenr. 89[IV,105]: 82. (1927)

O. E. Schulz: in H. G. A. Engler, Pflanzenr. 89[IV,105]: 94. (1927)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Draba carnosula

Draba longisquamosa

Draba carnosula

Draba longisquamosa