Draba cana |
Draba serpentina |
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Brewer's Draba, canescent Draba, canescent whitlow-mustard, cushion Draba, hoary Draba, hoary whitlow-grass, lance-leaf Draba, lance-leaf whitlow-grass |
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Habit | Perennials; caudex simple or branched (branches short); not scapose. | Perennials; (cespitose); caudex branched (with persistent leaves, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched or branched distally, (0.6–)1–3(–3.8) dm, pubescent throughout, trichomes simple, 0.5–1mm, with 4–10-rayed ones, 0.05–0.2 mm (mostly branched on basal parts). |
unbranched, 0.3–1.3 dm, glabrous throughout. |
Basal leaves | rosulate; petiolate; petiole ciliate throughout; blade linear to oblanceolate or oblong, (0.5–)0.8–2(–3.5) cm × 1.5–4(–11) mm, margins entire or dentate, (base and apex ciliate, trichomes simple, 0.3–0.8 mm), surfaces pubescent with short-stalked, 4–12-rayed trichomes, 0.1–0.3 mm. |
rosulate; sessile; blade obovate to obovate-oblanceolate, 0.4–1 cm × (2–)2.5–4(–5) mm, margins entire, (ciliate, trichomes simple, 0.2–0.5 mm, apex acute), surfaces glabrous. |
Cauline leaves | 3–10(–17), (not overlapping); sessile; blade lanceolate to ovate or oblong, margins entire or dentate, surfaces pubescent as basal (adaxially with simple and forked trichomes near blade base). |
0. |
Racemes | (10–)15–47(–63)-flowered, basally bracteate, often considerably elongated in fruit; rachis not flexuous, densely pubescent, trichomes 4–10-rayed (0.05–0.2 mm), and fewer simple ones. |
6–14-flowered, ebracteate, elongated in fruit; rachis not flexuous, glabrous. |
Flowers | sepals (green or lavender), oblong to ovate, 1.5–2 mm, pubescent, (trichomes simple and few-rayed); petals white, oblanceolate to spatulate, 2.3–3.7(–4.5) × 0.7–1.7 mm; anthers ovate, 0.1–0.2 mm. |
sepals ovate, 2–3 mm, glabrous; petals white, spatulate to obovate, 5–6.5 × 2–3 mm; anthers ovate to oblong, 0.4–0.5 mm. |
Fruiting pedicels | suberect or ascending, straight, 2–5(–10) mm, pubescent as rachis. |
divaricate-ascending, straight, 5–13 mm, glabrous. |
Fruits | (subappressed to rachis), linear-lanceolate to linear or, rarely, ovate-oblong, slightly twisted or plane, flattened, (5–)6–11 × 1.5–2(–2.5) mm; valves pubescent, trichomes short-stalked, 3–7-rayed, 0.05–0.3 mm; ovules 28–48 per ovary; style 0.1–0.6 mm. |
lanceolate to narrowly oblong, slightly twisted or plane, flattened, 6–12 × 2–3.5 mm; valves glabrous; ovules 8–16 per ovary; style 0.6–1.8(–2.5) mm. |
Seeds | ovoid, 0.5–0.7(–0.9) × 0.3–0.5 mm. |
oblong, 1.1–1.4 × 0.7–0.8 mm. |
2n | = 32. |
= 52. |
Draba cana |
Draba serpentina |
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Phenology | Flowering (May-)Jun–Aug. | Flowering May–Jul. |
Habitat | Rock outcrops and talus, open prairie benchlands, roadsides, meadows, alpine tundra | Rock outcrops, talus, and gravelly soil in mixed conifer and subalpine meadow communities |
Elevation | 0-4100 m (0-13500 ft) | 3200-3600 m (10500-11800 ft) |
Distribution |
AK; CA; CO; ID; ME; MI; MT; NH; NM; NV; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NF; NT; ON; QC; SK; YT; Greenland
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NV |
Discussion | The limits of Draba cana have long been confused, and the species was treated as a synonym of the Himalayan D. lanceolata Royle (M. L. Fernald 1934; C. L. Hitchcock 1941) or as a variety of the western North American D. breweri (R. C. Rollins 1993). However, G. A. Mulligan (1971) clearly demonstrated that all three are distinct and should be maintained. Some Utah plants corresponding to the type of D. valida have shorter and wider oblong-ovate fruits. In all other aspects, they are indistinguishable from D. cana. Additional studies are needed to establish whether such plants should be formally recognized. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Draba serpentina has been treated by other authors as a variety or synonym of D. oreibata. As indicated by I. A. Al-Shehbaz and M. D. Windham (2007), the two taxa are morphologically distinct, have different chromosome numbers, and are separated by over 480 kilometers. They are treated herein as separate species. Draba serpentina is known only from the Snake Range (White Pine County) and the Toiyabe Mountains (Lander County) in central Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 298. | FNA vol. 7, p. 335. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. breweri var. cana, D. valida | D. oreibata var. serpentina |
Name authority | Rydberg: Bull. Torrey Bot. Club 29: 241. (1902) | (Tiehm & P. K. Holmgren) Al-Shehbaz & Windham: Harvard Pap. Bot. 12: 414. (2007) |
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