Draba cana |
Draba santaquinensis |
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Brewer's Draba, canescent Draba, canescent whitlow-mustard, cushion Draba, hoary Draba, hoary whitlow-grass, lance-leaf Draba, lance-leaf whitlow-grass |
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Habit | Perennials; caudex simple or branched (branches short); not scapose. | Biennials or perennials; (short-lived); caudex simple (poorly developed, without persistent leaf bases); not scapose. |
Stems | unbranched or branched distally, (0.6–)1–3(–3.8) dm, pubescent throughout, trichomes simple, 0.5–1mm, with 4–10-rayed ones, 0.05–0.2 mm (mostly branched on basal parts). |
usually branched, 1.1–3.4 dm, hirsute proximally, trichomes mostly simple and 2-rayed, 0.2–1.5 mm, pubescent distally, trichomes mostly 2- or 3-rayed, 0.2–1 mm. |
Basal leaves | rosulate; petiolate; petiole ciliate throughout; blade linear to oblanceolate or oblong, (0.5–)0.8–2(–3.5) cm × 1.5–4(–11) mm, margins entire or dentate, (base and apex ciliate, trichomes simple, 0.3–0.8 mm), surfaces pubescent with short-stalked, 4–12-rayed trichomes, 0.1–0.3 mm. |
rosulate; petiole (obscure), ciliate, (trichomes simple and 2-rayed); blade obovate to oblanceolate, 1.6–2.5 cm × 5–10 mm, margins usually entire, rarely denticulate, surfaces pubescent, abaxially with stalked, mostly cruciform trichomes, 0.5–1 mm, adaxially with simple and stalked, 2–4-rayed ones. |
Cauline leaves | 3–10(–17), (not overlapping); sessile; blade lanceolate to ovate or oblong, margins entire or dentate, surfaces pubescent as basal (adaxially with simple and forked trichomes near blade base). |
usually 1–5; sessile or subsessile; blade oblanceolate to oblong, margins often denticulate, (ciliate proximally, with 2- or 3-rayed trichomes). |
Racemes | (10–)15–47(–63)-flowered, basally bracteate, often considerably elongated in fruit; rachis not flexuous, densely pubescent, trichomes 4–10-rayed (0.05–0.2 mm), and fewer simple ones. |
9–25-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent, trichomes 2- or 3-rayed. |
Flowers | sepals (green or lavender), oblong to ovate, 1.5–2 mm, pubescent, (trichomes simple and few-rayed); petals white, oblanceolate to spatulate, 2.3–3.7(–4.5) × 0.7–1.7 mm; anthers ovate, 0.1–0.2 mm. |
sepals oblong-obovate, 2–3.5 mm, pubescent, (trichomes short- to long-stalked, 2–4-rayed, 0.2–0.8 mm); petals yellow (often fading whitish), oblanceolate, 3–5 × 1–1.5 mm; anthers ovate, 0.4–0.5 mm. |
Fruiting pedicels | suberect or ascending, straight, 2–5(–10) mm, pubescent as rachis. |
horizontal to divaricate-ascending, usually straight, rarely curved upward, 7–10 mm, pubescent, trichomes 2- or 3-rayed. |
Fruits | (subappressed to rachis), linear-lanceolate to linear or, rarely, ovate-oblong, slightly twisted or plane, flattened, (5–)6–11 × 1.5–2(–2.5) mm; valves pubescent, trichomes short-stalked, 3–7-rayed, 0.05–0.3 mm; ovules 28–48 per ovary; style 0.1–0.6 mm. |
narrowly elliptic to slightly falcate, plane, flattened, (7–)10–16 × (2–)2.5–3.7 mm; valves pubescent, trichomes short-stalked, 2–4-rayed, 0.2–0.5 mm; ovules 14–22 per ovary; style 0.9–1.2 mm. |
Seeds | ovoid, 0.5–0.7(–0.9) × 0.3–0.5 mm. |
oblong, 1.1–1.5 × 0.7–0.9 mm. |
2n | = 32. |
= 20. |
Draba cana |
Draba santaquinensis |
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Phenology | Flowering (May-)Jun–Aug. | Flowering late Apr-early Jun. |
Habitat | Rock outcrops and talus, open prairie benchlands, roadsides, meadows, alpine tundra | Limestone outcrops and rocky slopes in mixed conifer communities |
Elevation | 0-4100 m (0-13500 ft) | 1800-2400 m (5900-7900 ft) |
Distribution |
AK; CA; CO; ID; ME; MI; MT; NH; NM; NV; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NF; NT; ON; QC; SK; YT; Greenland
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UT |
Discussion | The limits of Draba cana have long been confused, and the species was treated as a synonym of the Himalayan D. lanceolata Royle (M. L. Fernald 1934; C. L. Hitchcock 1941) or as a variety of the western North American D. breweri (R. C. Rollins 1993). However, G. A. Mulligan (1971) clearly demonstrated that all three are distinct and should be maintained. Some Utah plants corresponding to the type of D. valida have shorter and wider oblong-ovate fruits. In all other aspects, they are indistinguishable from D. cana. Additional studies are needed to establish whether such plants should be formally recognized. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although Draba santaquinensis was included within D. brachystylis by previous authors, I. A. Al-Shehbaz and M. D. Windham (2007) have shown that it is distinct both morphologically and chromosomally. It is currently known only from Utah County (American Fork, Provo, and Santaquin canyons) in north-central Utah. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 298. | FNA vol. 7, p. 334. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. breweri var. cana, D. valida | |
Name authority | Rydberg: Bull. Torrey Bot. Club 29: 241. (1902) | Windham & Allphin: Harvard Pap. Bot. 12: 410. (2007) |
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