Draba cana |
Draba crassifolia |
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Brewer's Draba, canescent Draba, canescent whitlow-mustard, cushion Draba, hoary Draba, hoary whitlow-grass, lance-leaf Draba, lance-leaf whitlow-grass |
Rocky Mountain Draba, Rocky Mountain whitlow-grass, snowbed Draba, snowbed whitlow-grass, thick-leaf Draba |
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Habit | Perennials; caudex simple or branched (branches short); not scapose. | Annuals or perennials; (short-lived); caudex branched (when present); usually scapose. |
Stems | unbranched or branched distally, (0.6–)1–3(–3.8) dm, pubescent throughout, trichomes simple, 0.5–1mm, with 4–10-rayed ones, 0.05–0.2 mm (mostly branched on basal parts). |
unbranched or, rarely, branched distally, (0.1–)0.3–1.1(–1.5) dm, usually glabrous throughout, rarely pubescent proximally, trichomes simple, 0.3–0.7 mm. |
Basal leaves | rosulate; petiolate; petiole ciliate throughout; blade linear to oblanceolate or oblong, (0.5–)0.8–2(–3.5) cm × 1.5–4(–11) mm, margins entire or dentate, (base and apex ciliate, trichomes simple, 0.3–0.8 mm), surfaces pubescent with short-stalked, 4–12-rayed trichomes, 0.1–0.3 mm. |
rosulate; petiolate; petiole ciliate throughout; blade oblanceolate to obovate, (0.2–) 0.5–2.5(–3) cm × (1–)2–4(–6) mm, margins usually entire, rarely denticulate, (sometimes ciliate), surfaces glabrous or sparsely pubescent, with simple and 2-rayed trichomes, 0.3–0.9 mm. |
Cauline leaves | 3–10(–17), (not overlapping); sessile; blade lanceolate to ovate or oblong, margins entire or dentate, surfaces pubescent as basal (adaxially with simple and forked trichomes near blade base). |
usually 0, rarely 1; sessile; blade oblong to ovate, margins entire, surfaces glabrous. |
Racemes | (10–)15–47(–63)-flowered, basally bracteate, often considerably elongated in fruit; rachis not flexuous, densely pubescent, trichomes 4–10-rayed (0.05–0.2 mm), and fewer simple ones. |
(2–)4–15(–25)-flowered, ebracteate, elongated in fruit; rachis slightly flexuous or straight, glabrous. |
Flowers | sepals (green or lavender), oblong to ovate, 1.5–2 mm, pubescent, (trichomes simple and few-rayed); petals white, oblanceolate to spatulate, 2.3–3.7(–4.5) × 0.7–1.7 mm; anthers ovate, 0.1–0.2 mm. |
(chasmogamous, petaliferous); sepals (green or purplish), ovate, 1–2 mm, glabrous; petals yellow (often fading white), oblanceolate, 1.5–2.5(–3) × 0.5–1 mm; anthers ovate, 0.15–0.25 mm. |
Fruiting pedicels | suberect or ascending, straight, 2–5(–10) mm, pubescent as rachis. |
horizontal to divaricate-ascending, usually straight, rarely curved upward, 3–8(–10) mm (subequaling or shorter than fruit), glabrous. |
Fruits | (subappressed to rachis), linear-lanceolate to linear or, rarely, ovate-oblong, slightly twisted or plane, flattened, (5–)6–11 × 1.5–2(–2.5) mm; valves pubescent, trichomes short-stalked, 3–7-rayed, 0.05–0.3 mm; ovules 28–48 per ovary; style 0.1–0.6 mm. |
usually narrowly elliptic to lanceolate, rarely linear-lanceolate, plane, flattened, (3–)5–10 × 1.5–2.5 mm; valves glabrous; ovules (8–)16–24(–30) per ovary; style 0.02–0.1 mm. |
Seeds | ovoid, 0.5–0.7(–0.9) × 0.3–0.5 mm. |
elliptic, 0.7–0.8 × 0.4–0.5 mm. |
2n | = 32. |
= 40. |
Draba cana |
Draba crassifolia |
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Phenology | Flowering (May-)Jun–Aug. | Flowering Jun–Sep. |
Habitat | Rock outcrops and talus, open prairie benchlands, roadsides, meadows, alpine tundra | Rock outcrops and talus, subalpine meadows, alpine summits and tundra, bare snow-melt areas |
Elevation | 0-4100 m (0-13500 ft) | (50-)1000-4300 m ((200-)3300-14100 ft) |
Distribution |
AK; CA; CO; ID; ME; MI; MT; NH; NM; NV; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NF; NT; ON; QC; SK; YT; Greenland
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AK; AZ; CO; ID; MT; NM; NV; UT; WA; WY; AB; BC; LB; NT; NU; QC; YT; Greenland; n Europe (Norway, Sweden)
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Discussion | The limits of Draba cana have long been confused, and the species was treated as a synonym of the Himalayan D. lanceolata Royle (M. L. Fernald 1934; C. L. Hitchcock 1941) or as a variety of the western North American D. breweri (R. C. Rollins 1993). However, G. A. Mulligan (1971) clearly demonstrated that all three are distinct and should be maintained. Some Utah plants corresponding to the type of D. valida have shorter and wider oblong-ovate fruits. In all other aspects, they are indistinguishable from D. cana. Additional studies are needed to establish whether such plants should be formally recognized. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
M. D. Windham (2004) presented morphological and chromosomal data suggesting that Draba crassifolia is an allopolyploid produced by hybridization between D. albertina and D. fladnizensis. Although the species is distinctive in large part, some individuals can be difficult to place and there is evidence of rare backcrossing (Windham, unpubl.). The attribution to Arizona is based on Schaack 345 (US) and Kearney & Peebles 12156 (US), both collected on the San Francisco Peaks in Coconino County. Draba crassifolia is found at elevations as low as 50 m in Greenland and the islands of Nunavut. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 298. | FNA vol. 7, p. 302. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. breweri var. cana, D. valida | D. crassifolia var. parryi, D. parryi |
Name authority | Rydberg: Bull. Torrey Bot. Club 29: 241. (1902) | Graham: Edinburgh New Philos. J. 7: 182. (1829) |
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