Draba breweri |
Draba palanderiana |
|
---|---|---|
Brewer's whitlow grass, cushion Draba |
palander's Draba, palander's whitlow-grass |
|
Habit | Perennials; (cespitose, grayish pubescent); caudex branched (sometimes covered with persistent leaf bases, branches short, compact); sometimes scapose. | Perennials; (cespitose); caudex branched (with persistent leaf remains, branches sometimes terminating in sterile rosettes); scapose. |
Stems | unbranched, (0.1–)0.2–1(–1.5) dm, pubescent throughout, trichomes stalked, 4–10-rayed, 0.1–0.3 mm. |
unbranched, 0.2–1.1(–1.5) dm, pubescent proximally, trichomes 2–9-rayed, 0.05–0.2 mm, glabrous or sparsely pubescent distally. |
Basal leaves | rosulate; petiolate; petiole base ciliate, margin not ciliate, (trichomes simple and 2-rayed, 0.3–0.8 mm); blade oblanceolate to obovate, (0.3–)0.4–1.5(–2.5) cm × 1.5–3(–5) mm, margins usually entire, rarely dentate, surfaces densely pubescent with stalked, 4–10-rayed, stellate trichomes, 0.1–0.2 mm (sometimes 1 or more rays spurred). |
rosulate; petiolate; petiole (thickened), ciliate proximally, (margin not ciliate, trichomes simple and 2-rayed, 0.2–0.5 mm); blade oblanceolate to obovate, 0.3–0.9(–1.5) cm × 1–4 mm, margins usually entire, rarely denticulate, surfaces pubescent with minutely stalked, 8–16-rayed, stellate trichomes, 0.08–0.3 mm, (midvein obscure abaxially). |
Cauline leaves | 0–3(–6); sessile; blade oblong or lanceolate to ovate, margins entire (sometimes ciliate at base), surfaces usually pubescent as basal. |
0. |
Racemes | (5–)7–18(–24)-flowered, ebracteate, slightly to considerably elongated in fruit; rachis not flexuous, pubescent with stalked, 4–10-rayed trichomes, (0.1–0.3 mm). |
5–17-flowered, ebracteate, elongated in fruit; rachis not flexuous, usually glabrous, rarely sparsely pubescent as stem. |
Flowers | sepals (persistent), ovate, 1.2–2 mm, pubescent, (trichomes short-stalked, 2–6-rayed); petals white, spatulate to oblanceolate, 2–3 × 0.7–1.1 mm; anthers ovate, 0.2–0.25 mm. |
sepals ovate, 2–2.5 mm, pubescent, (trichomes simple and short-stalked, 2–6-rayed); petals white or creamy white, obovate, 4.5–5.5 × 2–3 mm; anthers ovate, 0.4–0.6 mm. |
Fruiting pedicels | ascending, (sometimes slightly appressed to rachis), straight, 1.5–3(–4) mm, pubescent as rachis. |
divaricate-ascending, straight, (slender), 4–10 mm, usually glabrous, rarely sparsely pubescent as stem. |
Fruits | lanceolate or oblong to linear, usually slightly to strongly twisted, rarely plane, flattened, 3.5–9(–11) × 1.5–2.5 mm; valves pubescent, trichomes short-stalked, 2–5-rayed, 0.05–0.25 mm; ovules 28–40 per ovary; style 0.1–0.3(–0.4) mm. |
(often aborting, becoming lopsided), oblong to elliptic or ovate, plane, flattened, 4–8 × 2–3 mm; valves glabrous; ovules 18–32 per ovary; style 0.3 0.8 mm. |
Seeds | ovoid, 0.5–0.7 × 0.3–0.5 mm. |
ovoid, 0.9–1.1 × 0.5–0.7 mm. |
2n | = 32. |
= 16, 32, 64. |
Draba breweri |
Draba palanderiana |
|
Phenology | Flowering Jul–Aug. | Flowering Jun–Jul. |
Habitat | Rock outcrops, talus, exposed ridges, alpine areas | Rock outcrops, talus, Dryas fellfields, tundra |
Elevation | 3100-4100 m (10200-13500 ft) | 0-1800 m (0-5900 ft) |
Distribution |
CA
|
AK; NT; YT; e Asia (Russian Far East, Siberia) |
Discussion | The circumscription of Draba breweri was expanded by R. C. Rollins (1993) to include D. cana. Plants of D. cana differ from those of D. breweri by being non-cespitose (versus cespitose) and taller [(4–)10–30(–38) versus (1–)2–9(–15) cm], and by having basally bracteate (versus ebracteate) racemes, and stems, pedicels, and sepals pubescent with a mixture of simple and branched (versus exclusively branched) trichomes. Draba breweri is known to us from Alpine, Fresno, Inyo, Mono, Plumas, Tulare, and Tuolumne counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
North American plants of Draba palanderiana are diploid and appear to be self-incompatible, an unusual situation among arctic members of the genus. By contrast, some plants from the Russian Far East are tetraploid (2n = 32) or octoploid (2n = 64) and appear to be self-compatible. Detailed studies are needed to establish whether one or two taxa are involved. Many North American collections of Draba palanderiana have been misidentified as D. nivalis. The species is easily separated from D. nivalis by having ciliate (versus non-ciliate) bases of basal leaves, larger petals (4.5–5.5 × 2–3 versus 2–3.5 × 0.8–1.4 mm) that are pale yellow or cream (versus white), and longer fruiting pedicels 4–10 [versus 1–4.5(–8)] mm. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 297. | FNA vol. 7, p. 326. |
Parent taxa | Brassicaceae > tribe Arabideae > Draba | Brassicaceae > tribe Arabideae > Draba |
Sibling taxa | ||
Synonyms | D. breweri var. sublaxa | |
Name authority | S. Watson: Proc. Amer. Acad. Arts 23: 260. (1888) | Kjellman: in A. E. Nordenskiöld, Vega Exp. Vetensk. Iakttag. 2: 45. (1883) |
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