Draba brachystylis |
Draba ramulosa |
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shortstyle Draba |
Tushar Mountain Draba |
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Habit | Annuals or perennials; (short-lived); caudex often simple (poorly developed); not scapose. | Perennials; (loosely matted, grayish); caudex branched (with persistent leaf bases, branches creeping, sometimes terminating in sterile rosettes); not scapose. |
Stems | unbranched or branched, 0.6–3(–3.7) dm, pubescent throughout, trichomes simple, 0.4–0.9 mm, and 2–4-rayed, 0.05–0.4 mm. |
unbranched, 0.4–0.6 dm, densely pubescent throughout, trichomes dendritic, 3–6-rayed, (often crisped), 0.1–0.4 mm. |
Basal leaves | rosulate; subsessile; petiole base ciliate, (trichomes simple, 0.3–0.9 mm); blade oblanceolate, 1–3.5 cm × 4–7(–12) mm, margins entire or denticulate, surfaces pubescent, abaxially with stalked, (2–)4-rayed trichomes, 0.1–0.5 mm, adaxially with simple trichomes, 0.2–0.6 mm, with smaller, 2–4-rayed ones. |
(imbricate); not rosulate; sessile; blade obovate to oblanceolate, 0.4–1.1 cm × 2–3.2 mm, margins entire, (base and margins not ciliate), surfaces pubescent with stalked, 4–8-rayed trichomes, 0.1–0.5 mm, (sometimes 1 or more rays spurred), adaxially sometimes trichomes simple. |
Cauline leaves | (1 or) 2–6(–8); sessile; blade lanceolate or oblanceolate to ovate, margins entire or denticulate, surfaces pubescent as basal. |
2 or 3 (sometimes basal leaves spaced, flowering stem appearing to 8-leaved); sessile; blade oblong to ovate, margins entire, surfaces pubescent as basal. |
Racemes | (5–)10–35(–47)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
4–15-flowered, ebracteate or proximalmost 1 or 2 flowers bracteate, slightly elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals ovate, 2–2.7 mm, pubescent, (trichomes simple and short-stalked, 2-rayed); petals pale yellow (fading white), oblanceolate, 2.5–3.7 × 0.9–1.2 mm; anthers ovate, 0.2–0.3 mm. |
sepals broadly ovate, 1.7–2.4 mm, pubescent, (trichomes short-stalked, 2–5-rayed); petals yellow, obovate to oblanceolate, 3–4.5 × 1.5–2.5 mm; anthers ovate, 0.4–0.5 mm. |
Fruiting pedicels | divaricate-ascending, straight or only slightly curved upward, (1.5–)3–8(–10) mm, pubescent throughout, trichomes simple and stalked, 2–4-rayed. |
divaricate-ascending, usually straight, rarely curved upward, 3–6(–10) mm, pubescent, trichomes 3–6-rayed, (crisped, 0.1–0.4 mm), and, sometimes, simple. |
Fruits | narrowly elliptic to lanceolate, plane, flattened, (7–)10–16(–19) × 2–3.5 mm; valves pubescent, trichomes simple, spurred, and short-stalked, 2-rayed, 0.05–0.2(–0.3) mm; ovules 20–36(–42) per ovary; style 0.2–0.6(–0.8) mm. |
(not appressed to rachis), ovate to elliptic, plane, flattened, 4–6.5 × 2.5–4 mm; valves densely pubescent, trichomes simple and short-stalked, 2–5-rayed, 0.08–0.35 mm; ovules 6–12 per ovary; style (0.1–)0.3–0.7 mm. |
Seeds | oblong, 0.9–1.3 × 0.6–0.7 mm. |
oblong, 1.4–1.8 × 0.8–1.2 mm. |
2n | = 44. |
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Draba brachystylis |
Draba ramulosa |
|
Phenology | Flowering Jun–Jul. | Flowering Jun–Aug. |
Habitat | Fir and aspen communities, moist areas on rocky slopes | Rock outcrops, talus, gravelly soils |
Elevation | 1700-3000 m (5600-9800 ft) | 3300-3600 m (10800-11800 ft) |
Distribution |
UT |
UT |
Discussion | Of conservation concern. Based on morphological and chromosomal data (L. Allphin and M. D. Windham, unpubl.), Draba brachystylis is hypothesized to be an allopolyploid derived through hybridization between D. albertina and D. santaquinensis. It is known to us only from Cache, Duchesne, Juab, Salt Lake, and Utah counties. C. L. Hitchcock (1941) indicated that the species grows in the Charleston Mountains of Clark County, Nevada, but we have not seen unequivocal material of it from that area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Molecular and chromosomal data (M. D. Windham, unpubl.) strongly suggest that Draba ramulosa is an allopolyploid species. It is thought to have originated through hybridization between D. sobolifera and a member of the white-flowered, euploid lineage of M. A. Beilstein and M. D. Windham (2003). It is easily distinguished from D. sobolifera by having pale yellow to whitish (versus bright yellow) petals, grayish (versus green) foliage, non-ciliate (versus ciliate) basal leaves pubescent with 4–8-rayed (versus 2–4-rayed) trichomes, often proximally bracteate (versus ebracteate) racemes, and flattened (versus inflated basally) fruits. Draba ramulosa is known from the Tushar Mountains in south-central Utah (Beaver and Piute counties). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 297. | FNA vol. 7, p. 332. |
Parent taxa | ||
Sibling taxa | ||
Name authority | Rydberg: Bull. Torrey Bot. Club 29: 240. (1902) | Rollins: Contr. Gray Herb. 214: 6. (1984) |
Web links |