Draba brachystylis |
Draba pectinipila |
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shortstyle Draba |
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Habit | Annuals or perennials; (short-lived); caudex often simple (poorly developed); not scapose. | Perennials; (cespitose, sometimes forming mats); caudex branched (with persistent leaf bases, branches creeping, terminating in scapes or sterile rosettes); scapose. |
Stems | unbranched or branched, 0.6–3(–3.7) dm, pubescent throughout, trichomes simple, 0.4–0.9 mm, and 2–4-rayed, 0.05–0.4 mm. |
unbranched, (0.3–)0.4–1.6(–1.9) dm, pubescent, trichomes sessile, pectinate, 0.1–0.4 mm, (parallel to long axis of stem, sometimes with irregularly 2–4-rayed ones, 0.2–0.6 mm). |
Basal leaves | rosulate; subsessile; petiole base ciliate, (trichomes simple, 0.3–0.9 mm); blade oblanceolate, 1–3.5 cm × 4–7(–12) mm, margins entire or denticulate, surfaces pubescent, abaxially with stalked, (2–)4-rayed trichomes, 0.1–0.5 mm, adaxially with simple trichomes, 0.2–0.6 mm, with smaller, 2–4-rayed ones. |
rosulate; subsessile; blade narrowly oblanceolate to linear, (0.4–)0.6–1.3(–2) cm × 0.9–2.2 mm, margins entire (not ciliate), surfaces pubescent with subsessile or sessile, pectinate trichomes, 0.2–0.5 mm. |
Cauline leaves | (1 or) 2–6(–8); sessile; blade lanceolate or oblanceolate to ovate, margins entire or denticulate, surfaces pubescent as basal. |
0. |
Racemes | (5–)10–35(–47)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem. |
5–22-flowered, ebracteate, considerably elongated in fruit; rachis not flexuous, pubescent as stem. |
Flowers | sepals ovate, 2–2.7 mm, pubescent, (trichomes simple and short-stalked, 2-rayed); petals pale yellow (fading white), oblanceolate, 2.5–3.7 × 0.9–1.2 mm; anthers ovate, 0.2–0.3 mm. |
sepals broadly ovate, 2–3.2 mm, pubescent, (trichomes pectinate); petals yellow, obovate to spatulate, 4–6.5 × 1.5–2.5 mm; anthers ovate 0.4–0.5 mm. |
Fruiting pedicels | divaricate-ascending, straight or only slightly curved upward, (1.5–)3–8(–10) mm, pubescent throughout, trichomes simple and stalked, 2–4-rayed. |
divaricate-ascending, straight, (5–)7–14 mm, sparsely pubescent, trichomes pectinate. |
Fruits | narrowly elliptic to lanceolate, plane, flattened, (7–)10–16(–19) × 2–3.5 mm; valves pubescent, trichomes simple, spurred, and short-stalked, 2-rayed, 0.05–0.2(–0.3) mm; ovules 20–36(–42) per ovary; style 0.2–0.6(–0.8) mm. |
ovoid, plane, slightly inflated basally, 4–6(–7) × 2–3 mm; valves pubescent, trichomes usually sessile, pectinate, 0.2–0.5 mm, rarely with simple ones; ovules 4–8 per ovary; style 0.5–1.5 mm. |
Seeds | oblong, 0.9–1.3 × 0.6–0.7 mm. |
oblong, 1.1–1.5 × 0.6–0.8 mm. |
2n | = 44. |
= 22. |
Draba brachystylis |
Draba pectinipila |
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Phenology | Flowering Jun–Jul. | Flowering Apr–Jun. |
Habitat | Fir and aspen communities, moist areas on rocky slopes | Rocky slopes in sagebrush scrub and pinyon-juniper woodlands |
Elevation | 1700-3000 m (5600-9800 ft) | 1700-2400 m (5600-7900 ft) |
Distribution |
UT |
CO; MT; UT; WY |
Discussion | Of conservation concern. Based on morphological and chromosomal data (L. Allphin and M. D. Windham, unpubl.), Draba brachystylis is hypothesized to be an allopolyploid derived through hybridization between D. albertina and D. santaquinensis. It is known to us only from Cache, Duchesne, Juab, Salt Lake, and Utah counties. C. L. Hitchcock (1941) indicated that the species grows in the Charleston Mountains of Clark County, Nevada, but we have not seen unequivocal material of it from that area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Draba pectinipila was treated as a synonym of D. oligosperma by G. A. Mulligan (1972), R. C. Rollins (1993), and N. H. Holmgren (2005b). The latter recognized D. juniperina as a distinct species but overlooked the fact that it is indistinguishable from the type collections of the earlier-published D. pectinipila. The species (including D. juniperina) differs significantly from D. oligosperma in both chromosome number and morphology. Draba pectinipila is easily distinguished by having fruit valves pubescent with pectinate trichomes, fruiting pedicels (5–)7–14 mm, petals 4–6.5 mm, ovules 4–8 per ovary, and styles 0.5–1.5 mm. By contrast, D. oligosperma has fruit valves glabrous or pubescent with simple or 2-rayed trichomes, fruiting pedicels (2–)3–10(–13) mm, petals 2.5–4 mm, ovules 6–12 per ovary, and styles 0.1–0.8(–1.1) mm. Draba pectinipila was previously known only from the type locality in northwestern Wyoming (Park County). Its range is now expanded to include that of D. juniperina in northwestern Colorado (Moffat County), northeastern Utah (Daggett and Uintah counties), and southwestern Wyoming (Sweetwater County). The record from Uintah County is based on Goodrich 22275 (NY). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 297. | FNA vol. 7, p. 328. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | D. juniperina, D. oligosperma var. juniperina, D. oligosperma var. pectinipila | |
Name authority | Rydberg: Bull. Torrey Bot. Club 29: 240. (1902) | Rollins: Rhodora 55: 231. (1953) |
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