FNA: Draba brachystylis vs. Draba cyclomorpha

	Draba brachystylis	Draba cyclomorpha
	shortstyle Draba	granite Draba, Lemmon's Draba
Habit	Annuals or perennials; (short-lived); caudex often simple (poorly developed); not scapose.	Perennials; (cespitose, not pulvinate); caudex branched (with persistent petiole remains, branches sometimes terminating in sterile rosettes); scapose.
Stems	unbranched or branched, 0.6–3(–3.7) dm, pubescent throughout, trichomes simple, 0.4–0.9 mm, and 2–4-rayed, 0.05–0.4 mm.	unbranched, 0.1–0.6(–1) dm, (thinly) pilose throughout, trichomes simple and stalked, 2- (or 3-)rayed, (often crisped), 0.1–0.8 mm.
Basal leaves	rosulate; subsessile; petiole base ciliate, (trichomes simple, 0.3–0.9 mm); blade oblanceolate, 1–3.5 cm × 4–7(–12) mm, margins entire or denticulate, surfaces pubescent, abaxially with stalked, (2–)4-rayed trichomes, 0.1–0.5 mm, adaxially with simple trichomes, 0.2–0.6 mm, with smaller, 2–4-rayed ones.	rosulate; shortly petiolate; petiole base and margin ciliate, (trichomes simple, 0.2–1 mm); blade (somewhat fleshy), oblanceolate to obovate, 0.4–1 cm × 2–5 mm, margins entire, surfaces glabrous or hirsute, abaxially usually with stalked, 2-rayed trichomes, 0.1–0.6 mm, rarely with fewer, simple ones, (midvein obscure), adaxially with almost exclusively simple trichomes, to 0.6 mm.
Cauline leaves	(1 or) 2–6(–8); sessile; blade lanceolate or oblanceolate to ovate, margins entire or denticulate, surfaces pubescent as basal.	0.
Racemes	(5–)10–35(–47)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem.	5–14(–20)-flowered, ebracteate, elongated in fruit; rachis not flexuous, pubescent as stem.
Flowers	sepals ovate, 2–2.7 mm, pubescent, (trichomes simple and short-stalked, 2-rayed); petals pale yellow (fading white), oblanceolate, 2.5–3.7 × 0.9–1.2 mm; anthers ovate, 0.2–0.3 mm.	sepals ovate, 1.5–2.2 mm, pubescent, (trichomes simple and short-stalked, 2-rayed); petals yellow, spatulate, 4–5.5 × 1.5–2.5 mm; anthers oblong, 0.6–0.7 mm.
Fruiting pedicels	divaricate-ascending, straight or only slightly curved upward, (1.5–)3–8(–10) mm, pubescent throughout, trichomes simple and stalked, 2–4-rayed.	horizontal to divaricate-ascending, often curved upward or straight, 3–8(–10) mm, pilose as stem.
Fruits	narrowly elliptic to lanceolate, plane, flattened, (7–)10–16(–19) × 2–3.5 mm; valves pubescent, trichomes simple, spurred, and short-stalked, 2-rayed, 0.05–0.2(–0.3) mm; ovules 20–36(–42) per ovary; style 0.2–0.6(–0.8) mm.	oblong to oblong-ovate, plane, flattened, 3.5–9 × 2.5–4 mm; valves glabrous; ovules 8–12 per ovary; style 0.3–1 mm.
Seeds	oblong, 0.9–1.3 × 0.6–0.7 mm.	ovoid, 1.4–1.9 × 1–1.2 mm.
2n	= 44.

	Draba brachystylis	Draba cyclomorpha
Phenology	Flowering Jun–Jul.	Flowering Jul–Aug.
Habitat	Fir and aspen communities, moist areas on rocky slopes	Basaltic talus, stony ridges
Elevation	1700-3000 m (5600-9800 ft)	2500-3100 m (8200-10200 ft)
Distribution	UT [BONAP county map]	OR [BONAP county map]
Discussion	Of conservation concern. Based on morphological and chromosomal data (L. Allphin and M. D. Windham, unpubl.), Draba brachystylis is hypothesized to be an allopolyploid derived through hybridization between D. albertina and D. santaquinensis. It is known to us only from Cache, Duchesne, Juab, Salt Lake, and Utah counties. C. L. Hitchcock (1941) indicated that the species grows in the Charleston Mountains of Clark County, Nevada, but we have not seen unequivocal material of it from that area. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Draba cyclomorpha was treated by Schulz, C. L. Hitchcock (1941), and R. C. Rollins (1993) as a variety of D. lemmonii. The two taxa are quite distinct morphologically and separated by nearly 1600 kilometers, justifying their treatment as distinct species (I. A. Al-Shehbaz and M. D. Windham 2007). Draba cyclomorpha is known to us only from the Wallowa Mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 7, p. 297.	FNA vol. 7, p. 305.
Parent taxa	Brassicaceae > tribe Arabideae > Draba	Brassicaceae > tribe Arabideae > Draba
Sibling taxa	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis	D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis
Synonyms		D. lemmonii var. cyclomorpha
Name authority	Rydberg: Bull. Torrey Bot. Club 29: 240. (1902)	Payson: Amer. J. Bot. 4: 263. (1917)
Web links	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: OR Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Draba cyclomorpha

shortstyle Draba

granite Draba, Lemmon's Draba

Habit

Annuals or perennials; (short-lived); caudex often simple (poorly developed); not scapose.

Perennials; (cespitose, not pulvinate); caudex branched (with persistent petiole remains, branches sometimes terminating in sterile rosettes); scapose.

Stems

unbranched or branched, 0.6–3(–3.7) dm, pubescent throughout, trichomes simple, 0.4–0.9 mm, and 2–4-rayed, 0.05–0.4 mm.

unbranched, 0.1–0.6(–1) dm, (thinly) pilose throughout, trichomes simple and stalked, 2- (or 3-)rayed, (often crisped), 0.1–0.8 mm.

Basal leaves

rosulate;

subsessile;

petiole base ciliate, (trichomes simple, 0.3–0.9 mm);

blade oblanceolate, 1–3.5 cm × 4–7(–12) mm, margins entire or denticulate, surfaces pubescent, abaxially with stalked, (2–)4-rayed trichomes, 0.1–0.5 mm, adaxially with simple trichomes, 0.2–0.6 mm, with smaller, 2–4-rayed ones.

rosulate; shortly petiolate;

petiole base and margin ciliate, (trichomes simple, 0.2–1 mm);

blade (somewhat fleshy), oblanceolate to obovate, 0.4–1 cm × 2–5 mm, margins entire, surfaces glabrous or hirsute, abaxially usually with stalked, 2-rayed trichomes, 0.1–0.6 mm, rarely with fewer, simple ones, (midvein obscure), adaxially with almost exclusively simple trichomes, to 0.6 mm.

Cauline leaves

(1 or) 2–6(–8);

sessile;

blade lanceolate or oblanceolate to ovate, margins entire or denticulate, surfaces pubescent as basal.

Racemes

(5–)10–35(–47)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

5–14(–20)-flowered, ebracteate, elongated in fruit;

rachis not flexuous, pubescent as stem.

Flowers

sepals ovate, 2–2.7 mm, pubescent, (trichomes simple and short-stalked, 2-rayed);

petals pale yellow (fading white), oblanceolate, 2.5–3.7 × 0.9–1.2 mm;

anthers ovate, 0.2–0.3 mm.

sepals ovate, 1.5–2.2 mm, pubescent, (trichomes simple and short-stalked, 2-rayed);

petals yellow, spatulate, 4–5.5 × 1.5–2.5 mm;

anthers oblong, 0.6–0.7 mm.

Fruiting pedicels

divaricate-ascending, straight or only slightly curved upward, (1.5–)3–8(–10) mm, pubescent throughout, trichomes simple and stalked, 2–4-rayed.

horizontal to divaricate-ascending, often curved upward or straight, 3–8(–10) mm, pilose as stem.

Fruits

narrowly elliptic to lanceolate, plane, flattened, (7–)10–16(–19) × 2–3.5 mm;

valves pubescent, trichomes simple, spurred, and short-stalked, 2-rayed, 0.05–0.2(–0.3) mm;

ovules 20–36(–42) per ovary;

style 0.2–0.6(–0.8) mm.

oblong to oblong-ovate, plane, flattened, 3.5–9 × 2.5–4 mm;

valves glabrous;

ovules 8–12 per ovary;

style 0.3–1 mm.

Seeds

oblong, 0.9–1.3 × 0.6–0.7 mm.

ovoid, 1.4–1.9 × 1–1.2 mm.

= 44.

Draba brachystylis

Draba cyclomorpha

Phenology

Flowering Jun–Jul.

Flowering Jul–Aug.

Habitat

Fir and aspen communities, moist areas on rocky slopes

Basaltic talus, stony ridges

Elevation

1700-3000 m (5600-9800 ft)

2500-3100 m (8200-10200 ft)

Distribution

[BONAP county map]

Discussion

Of conservation concern.

Based on morphological and chromosomal data (L. Allphin and M. D. Windham, unpubl.), Draba brachystylis is hypothesized to be an allopolyploid derived through hybridization between D. albertina and D. santaquinensis. It is known to us only from Cache, Duchesne, Juab, Salt Lake, and Utah counties. C. L. Hitchcock (1941) indicated that the species grows in the Charleston Mountains of Clark County, Nevada, but we have not seen unequivocal material of it from that area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Draba cyclomorpha was treated by Schulz, C. L. Hitchcock (1941), and R. C. Rollins (1993) as a variety of D. lemmonii. The two taxa are quite distinct morphologically and separated by nearly 1600 kilometers, justifying their treatment as distinct species (I. A. Al-Shehbaz and M. D. Windham 2007). Draba cyclomorpha is known to us only from the Wallowa Mountains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 7, p. 297.

FNA vol. 7, p. 305.

Parent taxa

Brassicaceae > tribe Arabideae > Draba

Sibling taxa

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. cyclomorpha, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

D. abajoensis, D. albertina, D. aleutica, D. alpina, D. aprica, D. arabisans, D. arctica, D. arctogena, D. argyrea, D. arida, D. asprella, D. asterophora, D. aurea, D. aureola, D. bifurcata, D. borealis, D. brachycarpa, D. brachystylis, D. breweri, D. burkei, D. californica, D. cana, D. carnosula, D. chamissonis, D. cinerea, D. corrugata, D. corymbosa, D. crassa, D. crassifolia, D. cruciata, D. cuneifolia, D. cusickii, D. daviesiae, D. densifolia, D. exunguiculata, D. fladnizensis, D. glabella, D. globosa, D. graminea, D. grandis, D. grayana, D. heilii, D. helleriana, D. hitchcockii, D. howellii, D. incana, D. incerta, D. incrassata, D. inexpectata, D. jaegeri, D. juvenilis, D. kassii, D. kluanei, D. lactea, D. lemmonii, D. lonchocarpa, D. longisquamosa, D. macounii, D. maguirei, D. malpighiacea, D. micropetala, D. mogollonica, D. monoensis, D. mulliganii, D. murrayi, D. nemorosa, D. nivalis, D. norvegica, D. novolympica, D. oblongata, D. ogilviensis, D. oligosperma, D. oreibata, D. oxycarpa, D. palanderiana, D. pauciflora, D. paucifructa, D. paysonii, D. pectinipila, D. pedicellata, D. pennellii, D. petrophila, D. pilosa, D. platycarpa, D. porsildii, D. praealta, D. pterosperma, D. ramosissima, D. ramulosa, D. rectifructa, D. reptans, D. ruaxes, D. santaquinensis, D. saxosa, D. scotteri, D. serpentina, D. sharsmithii, D. sibirica, D. sierrae, D. simmonsii, D. smithii, D. sobolifera, D. spectabilis, D. sphaerocarpa, D. sphaeroides, D. standleyi, D. stenoloba, D. stenopetala, D. streptobrachia, D. streptocarpa, D. subalpina, D. subcapitata, D. subumbellata, D. trichocarpa, D. ventosa, D. verna, D. viridis, D. weberi, D. yukonensis, D. zionensis

Synonyms

D. lemmonii var. cyclomorpha

Name authority

Rydberg: Bull. Torrey Bot. Club 29: 240. (1902)

Payson: Amer. J. Bot. 4: 263. (1917)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora

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Draba brachystylis

Draba cyclomorpha

Draba brachystylis

Draba cyclomorpha